Anoectangium handelii

Schiffner

Ann. K. K. Naturhist. Hofmus. 27: 490, figs. 51–59. 1913,

Treatment appears in FNA Volume 27. Treatment on page 522. Mentioned on page 521, 537, 538.
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Leaves dense, hiding the stem, ligulate to short-elliptic or ovate, 0.4–0.5(–0.6) mm; apex broadly acute to rarely rounded, apiculus absent or of one cell; margins often 2-stratose in patches; costa usually ending (1–)3–4 cells before apex. Specialized asexual reproduction by ovate gemmae, sometimes uncommon, occasionally apiculate by a projection, of ca. 8 cells, born on rhizoids in leaf axils. Sporophytes unknown.

Habitat: Fissures in rock, calcareous and noncalcareous sandstone
Elevation: moderate to high elevations (900-1700 m)

Distribution

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Calif., Colo., Nev., s Europe, sw, c Asia.

Discussion

Anoectangium handelii is quite similar to Gymnostomum viridulum in habitat, size, general appearance, and production of axillary gemmae (a morphological phenocopy), but is immediately distinguished by its narrow adaxial costal groove. Though apparently rare (California in Inyo County, Colorado in Boulder and Larimer counties, Nevada in Clark County), its small size and similarity to sterile Gymnostomum make A. handelii easy for collectors to pass over. C. C. Heyn and I. Hernnstadt (2004b) provided additional discussion of this curious species.

Lower Taxa

No lower taxa listed.