Amer. J. Sci. Arts 42: 49. 1842.
Biennials or, rarely, perennials; (short-lived, with caudex); usually densely hirsute (at least basally), trichomes simple (to 1 mm), sometimes mixed with stalked, forked ones. Stems simple or several from base (rosette), erect, often branched distally, (1.8–)2.5–4.5(–5.5) dm, (hirsute at least basally). Basal leaves: petiole (0.7–)2–5 cm; blade usually ovate to oblanceolate, rarely cordate, (0.8–)1.5–3 cm × 5–17 mm, margins dentate or serrate, surfaces sparsely to densely pubescent, trichomes simple or forked. Cauline leaves (5–)8–20(–27), (overlapping or not); blade ovate to oblong or lanceolate, (2–)3–7.5(–9) cm × (10–)13–25(–35) mm, base amplexicaul, margins usually dentate or serrate, sometimes entire, surfaces hirsute. Racemes branched. Fruiting pedicels ascending to divaricate-ascending, (7–)10–20(–24) mm, (glabrous or sparsely pubescent). Flowers: sepals oblong, 2.5–4.5 × 1–1.5 mm, lateral pair slightly saccate basally; petals white, spatulate, (5–)6–9 × 2–3.5 mm, apex obtuse; filaments 3–5 mm; anthers oblong, 0.8–1.2 mm. Fruits divaricate to erect-ascending, (not appressed to rachis), torulose, 2.2–4.6 cm × 0.8–1.2mm; valves each with prominent midvein extending to middle or full length; ovules 16–28 per ovary; style 0.5–2 mm, (slender). Seeds narrowly winged distally, oblong, 1–1.6 × 0.6–0.8 mm; wing 0.05–0.1 mm.
Phenology: Flowering Apr–May.
Habitat: Rocky wooded slopes, shady stream banks, limestone ledges and bluffs
Elevation: 0-500 m
D.C., Ind., Ky., Md., N.C., Ohio, Pa., Tenn., Va., W.Va.