Enum. Pl. Javae 2: 241. 1828. 1828
Plants terrestrial. Stems moderately long- to short-creeping, stolons absent. Leaves monomorphic, evergreen. Petiole ± as long as blade, base not swollen; vascular bundles more than 3, arranged in an arc, ± round in cross section. Blade broadly deltate or pentagonal, 2–3-pinnate-pinnatifid, gradually to abruptly reduced distally to pinnate or pinnatifid apex, papery to somewhat leathery. Pinnae not articulate to rachis, segment margins and especially apex spinulose; proximal pinnae largest, petiolulate, inequilateral with basal basiscopic pinnule much larger and more elongate than more distal pinnules; costae adaxially grooved, grooves continuous from rachis to costae to costules; indument of hairlike scales abaxially, absent adaxially. Veins free, forked. Sori in 1 row between midrib and margin, round; indusia round-reniform, attached at narrow sinus, persistent. Spores brownish, rugate or tuberculate, sometimes spiny. x = 41.
Tropics and subtropics, mostly in e Asia and Pacific Islands, a few in Africa, ca. 4 in Mexico, Central America, South America.
Species ca. 50 (1 in the flora naturalized from Asia).