Dicranostegia

(A. Gray) Pennell

Proc. Acad. Nat. Sci. Philadelphia 99: 189. 1947.

Etymology: Greek dicranos, two-headed, and stegos, sheath or cover, alluding to two-lobed calyx
Basionym: Cordylanthus Nuttall ex Bentham [unranked] Dicranostegia A. Gray Proc. Amer. Acad. Arts 19: 95. 1883
Treatment appears in FNA Volume 17. Treatment on page 679. Mentioned on page 458, 666, 670.

Herbs, annual; hemiparasitic. Stems erect or ascending, rarely prostrate, not fleshy, pubescent, hirsute, or glabrous. Leaves cauline, alternate; petiole absent; blade not fleshy, not leathery, margins pinnately 8–11-lobed. Inflorescences terminal, spikes; bracts present. Pedicels absent; bracteoles absent. Flowers: sepals 2, calyx bilaterally symmetric, spathelike, lobes triangular; petals 5, corolla white to yellow, marked with purple, strongly bilabiate, club-shaped, often gibbous at base of lobes, abaxial lobes 3, adaxial lobes 2, adaxial lip galeate; stamens 2, filaments glabrous or proximally pubescent, pollen sacs separate, connective elongate; staminodes 2, filiform; ovary 2-locular, placentation axile; stigma not expanded. Capsules: dehiscence loculicidal. Seeds 5–20, brown, ovoid, wings absent. x = 8.

Distribution

Calif., nw Mexico.

Discussion

Species 1.

Dicranostegia is sometimes classified as Cordylanthus subg. Dicranostegia (A. Gray) T. I. Chuang & Heckard. D. C. Tank et al. (2009) have shown that Dicranostegia is closely related to Chloropyron and that both genera form a clade that is sister to Castilleja and Triphysaria.

Like Chloropyron and Cordylanthus, Dicranostegia has closed, club-shaped flowers with a spathelike calyx. Flowers are subtended by a single, elongate bracteole and pinnate bracts. Dicranostegia differs in having pinnately lobed leaves, deeply two-parted calyces that are about half as long as the corollas, and stamens with pollen sacs held apart on relatively short connectives.