Dryopteris intermedia

(Muhlenberg ex Willdenow) A. Gray

Manual 630. 1848

Common names: Evergreen wood fern fancy fern dryoptère spinuleuse
Basionyms: Aspidium intermedium Muhlenberg ex Willdenow Sp. Pl. 5(1): 262. 1810
Synonyms: Dryopteris austriaca var. intermedia (Muhlenberg ex Willdenow) C. V. Morton Dryopteris spinulosa var. intermedia (Muhlenberg ex Willdenow) L. Underwood
Treatment appears in FNA Volume 2.
Click plate for higher resolution version.
Illustrator: John Myers
Leaves monomorphic, green through winter, 32–90 × 10–20 cm. Petiole 1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade green, ovate, 3-pinnate-pinnatifid, herbaceous, glandular. Pinnae ± in plane of blade, lanceolate-oblong; basal pinnae lanceolate, not reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. Sori midway between midvein and margin of segments. Indusia with minute glandular hairs. 2n = 82.

Habitat: Moist rocky woods, especially hemlock hardwoods, ravines, and edges of swamps
Elevation: 0–2000 m


V2 508-distribution-map.gif

N.B., Nfld., N.S., Ont., P.E.I., Que., Ala., Conn., Del., Ga., Ill., Ind., Iowa, Ky., Maine, Md., Mass., Mich., Minn., Mo., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., Tenn., Vt., Va., W.Va., Wis.


A related taxon, Dryopteris intermedia subsp. maderensis (J. Milde ex Alston) Fraser-Jenkins, occurs on eastern Atlantic islands.

Dryopteris intermedia and the other taxa in the " D. spinulosa complex" have long confounded taxonomists. Dryopteris intermedia is diploid and is one of the parents of the allotetraploids D. carthusiana and D. campyloptera. Dryopteris intermedia hybridizes with eight species. All hybrids are easily detected by the distinctive glandular hairs on the indusia and, usually, on the costae and costules.



Lower Taxa

No lower taxa listed.