Hylocomiastrum umbratum

(Hedwig) M. Fleischer ex Brotherus in H. G. A. Engler et al.

in H. G. A. Engler et al., Nat. Pflanzenfam. ed. 2, 11: 487. 1925

Illustrated
Basionyms: Hypnum umbratum Hedwig Sp. Musc. Frond., 263, plate 67, figs. 10 – 13. 1801
Synonyms: Hylocomium umbratum (Hedwig) Schimper
Treatment appears in FNA Volume 28. Treatment on page 327.
Plants to 15 cm, wiry. Stems 1–2 mm wide across leafy stem, irregularly to regularly 1–2(–3)-pinnate, often somewhat frondose or bushy, branches slender, innovations arching. Stem leaves erect- to wide-spreading, distant, broadly ovate to deltoid, slightly concave, 0.8–2(–2.5) × 0.8–1.5 mm; margins serrulate basally, spinose-serrate distally; apex gradually narrowed; costa usually double, sometimes single, single and 2-fid, or triple; medial laminal cells 30–65 × 4–7 µm. Branch leaves loosely imbricate, ovate, 0.4–1.2 × 0.3–0.8 mm; costa often single, but variable. Seta 2–4.5 cm. Capsule ovoid-ellipsoid, 1.5–2.5 mm.

Habitat: Soil, humus, old logs, and rock in moist forests, especially under spruce and fir
Elevation: low to high elevations (0-2000 m)

Distribution

V28 509-distribution-map.gif

B.C., N.B., Nfld. and Labr., N.S., Ont., P.E.I., Que., Alaska, Maine, Mass., Mich., N.H., N.Y., N.C., Tenn., Vt., Eurasia, Africa.

Discussion

In addition to the numerous highly branched paraphyllia, stems in Hylocomiastrum umbratum (but not H. pyrenaicum) also bear some scattered larger paraphyllia. These are more than twice the size of typical paraphyllia, are lanceolate-subulate with toothed margins, and closely resemble the foliose pseudoparaphyllia. Under a dissecting microscope these larger paraphyllia are clearly visible at 30× scattered among the shorter paraphyllia. In otherwise difficult specimens, the presence of these larger foliose paraphyllia can be used to distinguish H. umbratum from H. pyrenaicum.

Selected References

None.

Lower Taxa

None.

AuthorJoseph R. Rohrer +
Authority(Hedwig) M. Fleischer ex Brotherus in H. G. A. Engler et al. +
BasionymsHypnum umbratum +
DistributionB.C. +, N.B. +, Nfld. and Labr. +, N.S. +, Ont. +, P.E.I. +, Que. +, Alaska +, Maine +, Mass. +, Mich. +, N.H. +, N.Y. +, N.C. +, Tenn. +, Vt. +, Eurasia + and Africa. +
Elevationlow to high elevations (0-2000 m) +
HabitatSoil, humus, old logs, and rock in moist forests, especially under spruce and fir +
IllustrationPresent +
Illustration copyrightFlora of North America Association +
IllustratorPatricia M. Eckel +
Publication titlein H. G. A. Engler et al., Nat. Pflanzenfam. ed. +
Publication year1925 +
ReferenceNone +
Source xmlhttps://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f50eec43f223ca0e34566be0b046453a0960e173/coarse grained fna xml/V28/V28 509.xml +
Special statusIllustrated +
SynonymsHylocomium umbratum +
Taxon familyHylocomiaceae +
Taxon nameHylocomiastrum umbratum +
Taxon parentHylocomiastrum +
Taxon rankspecies +
VolumeVolume 28 +