Physaria gracilis

(Hooker) O’Kane & Al-Shehbaz

Novon 12: 323. 2002.

Common names: Spreading bladderpod
Basionym: Vesicaria gracilis Hooker Bot. Mag. 63: plate 3533. 1836
Synonyms: Alyssum gracile (Hooker) Kuntze Lesquerella gracilis (Hooker) S. Watson
Treatment appears in FNA Volume 7. Treatment on page 641. Mentioned on page 619.

Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). Stems simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. Basal leaves: blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). Cauline leaves: (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. Racemes loose, (elongated). Fruiting pedicels (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). Flowers: sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). Fruits (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. Seeds slightly flattened.


V7 1063-distribution-map.gif

Ala., Ill., Iowa, Kans., La., Miss., Mo., Okla., Tenn., Tex.


Subspecies 2 (2 in the flora).

Selected References



1 Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded. Physaria gracilis subsp. gracilis
1 Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate. Physaria gracilis subsp. nuttallii