Proc. Amer. Acad. Arts 34: 541. 1899.
Plants 30–70+ cm (perennating bases ± erect, internodes not winged). Leaves all or mostly opposite (distal sometimes alternate); blades ovate-deltate or ovate to lance-ovate, 2–6 × 1–5 cm, bases rounded to cuneate, margins toothed to entire, apices obtuse to acute, faces ± scabrellous. Heads borne singly. Involucres ± hemispheric, 15–22+ mm diam. Phyllaries 24–30+ in 3–4 series, ± erect, oblong, orbiculate, or ovate, 5–12+ mm. Ray florets ± 13; laminae 25 mm. Disc florets 80–120+; corollas yellow. Cypselae dark brown, narrowly elliptic, 7 mm, faces glabrous; pappi 0–0.5 mm. 2n = 34.
Phenology: Flowering (May–)Sep–Nov.
Habitat: Shaded slopes, limestone, juniper scrublands
Elevation: 100–500 m