Treatment appears in FNA Volume 28. Treatment on page 629. Mentioned on page 341, 342, 647, 475, 637.
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Plants small to large, in dense or loose mats, glaucous, green, brown, or yellowish brown, dull. Stems creeping, sparsely to profusely branched, irregularly pinnate; paraphyllia absent. Stem and branch leaves differentiated. Stem leaves scalelike, minute; apex acute-acuminate to rounded; costa single, long, ending below apex, thick, usually pellucid, or double and short; laminal cells short. Branch leaves with costa single, ending sharply at or near apex, pellucid. Sexual condition dioicous. Seta dark to light reddish or brown, flexuose. Capsule erect, exserted; operculum conic to obliquely short-rostrate; exostome whitish yellow to pale brown, often striolate at base and papillose.


North America, Mexico, West Indies, Central America, South America, Europe, e Asia, Africa, Pacific Islands, Australia, circumboreal areas.


Genera 4, species ca. 20 (2 genera, 9 species in the flora).

Lower Taxa


1 Branch leaves broadly ovate to lanceolate, ± abruptly narrowed mid leaf; laminal cells with papillae 1 or many. Anomodon
1 Branch leaves ovate-lanceolate, gradually tapered from base to apex; laminal cells smooth. Herpetineuron