Asteraceae (tribe Heliantheae) subtribe Coreopsidinae
Linnaea 5: 153. 1830
Mostly subtropical and warm-temperate New World and Old World.
Genera 20, species 374 (7 genera, 69 species in the flora).
The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle.
In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit.
|1||Phyllaries (excluding calyculi) 3–6 in 1(–2) series; disc florets 3–4+ (functionally staminate)||Dicranocarpus|
|1||Phyllaries (3–)8–34+ in ± 2 series; disc florets 3–10, 10–20, or (5–)12–150+ (bisexual, fertile)||> 2|
|2||Phyllaries connate 1/5–7/8+ their lengths||Thelesperma|
|2||Phyllaries usually distinct, rarely connate ± 1/10 their lengths||> 3|
|3||Cypselae (at least inner) ± 4-angled, ± linear-fusiform, often apically attenuate or beaked (none winged)||> 4|
|3||Cypselae all ± obcompressed (sometimes winged)||> 5|
|4||Disc florets 10–20 (staminal filaments hairy near anthers); cypselae usually with 1 groove on each face||Cosmos|
|4||Disc florets (5–)12–150+ (staminal filaments not hairy); cypselae with 0 or 2grooves on each face||Bidens|
|5||Annuals; ray florets 1–3 (laminae 1–2+ mm); cypselae mostly ellipsoid orobovoid (inner obovoid to obscurely urceolate, ± beaked)||Heterosperma|
|5||Annuals, perennials, subshrubs, or shrubs; ray florets usually 1, 2, 3, 5, 8, 13, or 21+, sometimes 0 (laminae mostly 4–30+ mm); cypselae mostly cuneate, linear, oblanceolate, oblong, ± orbiculate, or ovate (not beaked).||> 6|
|6||Cypselae rarely winged (margins sometimes thickened, winged in B. aristosa and B. polylepis); pappi usually of barbellate (rarely smooth) awns, sometimes 0||Bidens|
|6||Cypselae (some or all) usually thin-margined or ± winged; pappi usually 0, sometimes coroniform, or of 2 bristly cusps or scales (in Coreopsis), or of 1–2 retrorsely barbellate awns (in Coreocarpus)||> 7|
|7||Ray florets usually neuter or styliferous and sterile; wings of cypselae membranous, chartaceous, or corky, entire or lobed to toothed, sometimes ciliate||Coreopsis|
|7||Ray florets usually pistillate and fertile; wings of cypselae ± corky, ± pectinately toothed||Coreocarpus|