Difference between revisions of "Cheilanthes villosa"

Davenport ex Maxon

Proc. Biol. Soc. Wash. 31: 142. 1918.

Common names: Villous lip fern
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem diameter"><b>Stems </b>compact, 4–8 mm diam.;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale size or width;central stripe width;central stripe prominence;central stripe coloration;margin coloration;margin shape;margin course;margin arrangement or shape;margin fixation or orientation;margin duration">scales mostly bicolored, with broad, well-defined, dark, central stripe and narrow, light-brown margins, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Leaves </b>clustered, 7–30 cm;</span> <span class="statement" id="st-d0_s3" data-properties="leaf arrangement or growth form;leaf some measurement">vernation noncircinate.</span> <span class="statement" id="st-d0_s4" data-properties="petiole coloration;petiole shape"><b>Petiole </b>usually dark-brown, rounded adaxially.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade architecture or shape;blade width"><b>Blade </b>oblong-lanceolate to ovate, 3–4-pinnate at base, 1.5–5 cm wide;</span> <span class="statement" id="st-d0_s6" data-properties="rachis shape;rachis pubescence;scale shape;scale shape;scale shape;scale shape">rachis rounded adaxially, with scattered filiform to lanceolate scales, not pubescent.</span> <span class="statement" id="st-d0_s7" data-properties="pinna architecture;pinna coloration;basal pinna size;basal pinna architecture or shape;basal pinna pubescence"><b>Pinnae </b>not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing villous adaxially.</span> <span class="statement" id="st-d0_s8" data-properties="costa length"><b>Costae </b>green adaxially for most of length;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial scale architecture or arrangement;abaxial scale shape;abaxial scale shape;abaxial scale shape;abaxial scale shape;abaxial scale prominence;abaxial scale size;abaxial scale width;abaxial scale arrangement;basal lobe arrangement;ultimate segment position;ultimate segment architecture or shape;ultimate segment architecture or pubescence or shape">abaxial scales multiseriate, ovate to lanceolate, shallowly cordate at base, often with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, often concealing ultimate segments, erose-dentate, not ciliate.</span> <span class="statement" id="st-d0_s10" data-properties="ultimate segment shape;ultimate segment shape;ultimate segment shape;ultimate segment shape;ultimate segment size;ultimate segment some measurement;ultimate segment pubescence;ultimate segment pubescence;hair count;hair relief;hair relief;hair architecture"><b>Ultimate </b>segments round to oval, beadlike, the largest 1–2 mm, abaxially nearly glabrous except for a few coarse hairs, adaxially villous with coarse, unbranched hairs.</span> <span class="statement" id="st-d0_s11" data-properties="false indusium position;false indusium width"><b>False </b>indusia marginal to obscurely inframarginal, slightly differentiated, 0.05–0.25 mm wide.</span> <span class="statement" id="st-d0_s12" data-properties="sorus architecture"><b>Sori </b>± continuous around segment margins.</span> <span class="statement" id="st-d0_s13" data-properties="spore count"><b>Sporangia </b>containing 32 spores.</span> <span class="statement" id="st-d0_s14" data-properties="n=2n chromosome count">n = 2n = 90, apogamous.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, 4–8 mm diam.; scales mostly bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. <b>Leaves</b> clustered, 7–30 cm; vernation noncircinate. <b>Petiole</b> usually dark brown, rounded adaxially. <b>Blade</b> oblong-lanceolate to ovate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, with scattered filiform to lanceolate scales, not pubescent. <b>Pinnae</b> not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing villous adaxially. <b>Costae</b> green adaxially for most of length; abaxial scales multiseriate, ovate to lanceolate, shallowly cordate at base, often with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, often concealing ultimate segments, erose-dentate, not ciliate. <b>Ultimate</b> segments round to oval, beadlike, the largest 1–2 mm, abaxially nearly glabrous except for a few coarse hairs, adaxially villous with coarse, unbranched hairs. <b>False</b> indusia marginal to obscurely inframarginal, slightly differentiated, 0.05–0.25 mm wide. <b>Sori</b> ± continuous around segment margins. <b>Sporangia</b> containing 32 spores. n = 2n = 90, apogamous.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on limestone
 
|habitat=Cliffs and rocky slopes, usually on limestone
 
|elevation=400–2200 m
 
|elevation=400–2200 m
 
|distribution=Ariz.;N.Mex.;Tex.;n Mexico.
 
|distribution=Ariz.;N.Mex.;Tex.;n Mexico.
|discussion=<p>Cheilanthes villosa is an apogamous triploid of unknown parentage. Although there are reports of hybridization between C. villosa and C. eatonii (D. B. Lellinger 1985), recent gene exchange is unlikely because both taxa are apogamous in North America. Morphologically intermediate specimens (tentatively called C. pinkavii in T. Reeves 1979) are included here in C. eatonii, pending further study and valid publication of Reeve's epithet.</p>
+
|discussion=<p><i>Cheilanthes villosa</i> is an apogamous triploid of unknown parentage. Although there are reports of hybridization between <i>C. villosa</i> and <i>C. eatonii</i> (D. B. Lellinger 1985), recent gene exchange is unlikely because both taxa are apogamous in North America. Morphologically intermediate specimens (tentatively called C. pinkavii in T. Reeves 1979) are included here in <i>C. eatonii</i>, pending further study and valid publication of Reeve's epithet.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Cheilanthes villosa
 
name=Cheilanthes villosa
|author=
 
 
|authority=Davenport ex Maxon
 
|authority=Davenport ex Maxon
 
|rank=species
 
|rank=species
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|basionyms=
 
|basionyms=
 
|family=Pteridaceae
 
|family=Pteridaceae
 +
|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on limestone
 
|habitat=Cliffs and rocky slopes, usually on limestone
 
|elevation=400–2200 m
 
|elevation=400–2200 m
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|publication year=1918
 
|publication year=1918
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_816.xml
+
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-data-curation.git/src/bb6b7e3a7de7d3b7888a1ad48c7fd8f5c722d8d6/coarse_grained_fna_xml/V2/V2_816.xml
 
|genus=Cheilanthes
 
|genus=Cheilanthes
 
|species=Cheilanthes villosa
 
|species=Cheilanthes villosa
|abaxial scale architecture or arrangement=multiseriate
 
|abaxial scale arrangement=imbricate
 
|abaxial scale prominence=conspicuous
 
|abaxial scale shape=cordate;ovate;lanceolate
 
|abaxial scale size=largest
 
|abaxial scale width=0.4mm;1.5mm
 
|basal lobe arrangement=overlapping
 
|basal pinna architecture or shape=equilateral
 
|basal pinna pubescence=villous
 
|basal pinna size=not conspicuously larger than adjacent pair
 
|blade architecture or shape=3-4-pinnate
 
|blade shape=oblong-lanceolate;ovate
 
|blade width=1.5cm;5cm
 
|central stripe coloration=dark
 
|central stripe prominence=well-defined
 
|central stripe width=broad
 
|costa length=green
 
|false indusium position=marginal
 
|false indusium width=wide
 
|hair architecture=unbranched
 
|hair count=few
 
|hair relief=coarse;coarse
 
|leaf arrangement or growth form=clustered
 
|leaf some measurement=7cm;30cm
 
|margin arrangement or shape=contorted
 
|margin coloration=light-brown
 
|margin course=straight
 
|margin duration=persistent
 
|margin fixation or orientation=appressed
 
|margin shape=linear-lanceolate
 
|n=2n chromosome count=90
 
|petiole coloration=dark-brown
 
|petiole shape=rounded
 
|pinna architecture=not articulate
 
|pinna coloration=dark color
 
|rachis pubescence=not pubescent
 
|rachis shape=rounded
 
|scale coloration=bicolored
 
|scale shape=filiform;lanceolate
 
|scale size or width=narrow
 
|sorus architecture=continuous
 
|spore count=32
 
|stem architecture or arrangement=compact
 
|stem diameter=4mm;8mm
 
|ultimate segment architecture or pubescence or shape=not ciliate
 
|ultimate segment architecture or shape=erose-dentate
 
|ultimate segment position=concealing
 
|ultimate segment pubescence=villous;glabrous
 
|ultimate segment shape=beadlike;round;oval
 
|ultimate segment size=largest
 
|ultimate segment some measurement=1mm;2mm
 
 
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-->[[Category:Treatment]][[Category:Cheilanthes]]
 
-->[[Category:Treatment]][[Category:Cheilanthes]]

Revision as of 22:15, 27 May 2020

Stems compact, 4–8 mm diam.; scales mostly bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. Leaves clustered, 7–30 cm; vernation noncircinate. Petiole usually dark brown, rounded adaxially. Blade oblong-lanceolate to ovate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, with scattered filiform to lanceolate scales, not pubescent. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing villous adaxially. Costae green adaxially for most of length; abaxial scales multiseriate, ovate to lanceolate, shallowly cordate at base, often with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, often concealing ultimate segments, erose-dentate, not ciliate. Ultimate segments round to oval, beadlike, the largest 1–2 mm, abaxially nearly glabrous except for a few coarse hairs, adaxially villous with coarse, unbranched hairs. False indusia marginal to obscurely inframarginal, slightly differentiated, 0.05–0.25 mm wide. Sori ± continuous around segment margins. Sporangia containing 32 spores. n = 2n = 90, apogamous.


Phenology: Sporulating summer–fall.
Habitat: Cliffs and rocky slopes, usually on limestone
Elevation: 400–2200 m

Distribution

V2 816-distribution-map.gif

Ariz., N.Mex., Tex., n Mexico.

Discussion

Cheilanthes villosa is an apogamous triploid of unknown parentage. Although there are reports of hybridization between C. villosa and C. eatonii (D. B. Lellinger 1985), recent gene exchange is unlikely because both taxa are apogamous in North America. Morphologically intermediate specimens (tentatively called C. pinkavii in T. Reeves 1979) are included here in C. eatonii, pending further study and valid publication of Reeve's epithet.

Selected References

None.

Lower Taxa

None.