Difference between revisions of "Myricaceae"

Blume
Common names: Wax-myrtle Family
Treatment appears in FNA Volume 3. Treatment on page 429.
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--><span class="statement" id="st-d0_s0" data-properties="shrub duration;shrub duration;shrub odor;shrub coating;tree size;tree duration;tree duration;tree odor;tree coating"><b>Shrubs </b>or small trees, evergreen or deciduous, usually aromatic and resinous.</span> <span class="statement" id="st-d0_s1" data-properties=""><b>Roots </b>commonly with nitrogen-fixing nodules.</span> <span class="statement" id="st-d0_s2" data-properties="leaf arrangement;leaf architecture;leaf shape"><b>Leaves </b>alternate, simple or pinnatifid;</span> <span class="statement" id="st-d0_s3" data-properties="stipule presence;stipule presence">stipules absent or present;</span> <span class="statement" id="st-d0_s4" data-properties="petiole presence">petiole present.</span> <span class="statement" id="st-d0_s5" data-properties="glandular trichome architecture;glandular trichome architecture"><b>Leaf-</b>blade commonly with peltate, multicellular, glandular trichomes.</span> <span class="statement" id="st-d0_s6" data-properties=""><b>Inflorescences </b>axillary catkins;</span> <span class="statement" id="st-d0_s7" data-properties="beak presence">bracts present.</span> <span class="statement" id="st-d0_s8" data-properties="flower reproduction;flower reproduction;flower architecture;flower architecture"><b>Flowers </b>usually unisexual, occasionally bisexual, staminate and pistillate flowers usually on different plants, occasionally on same plants;</span> <span class="statement" id="st-d0_s9" data-properties="perianth presence">perianth absent.</span> <span class="statement" id="st-d0_s10" data-properties="flower architecture;bract architecture or arrangement or growth form"><b>Staminate </b>flowers subtended by solitary bract;</span> <span class="statement" id="st-d0_s11" data-properties="stamen atypical quantity;stamen quantity;stamen position;stamen position">stamens 2-14 (-22), hypogynous or ± epigynous;</span> <span class="statement" id="st-d0_s12" data-properties="filament shape;filament fusion;filament fusion">filaments filiform, distinct or basally connate;</span> <span class="statement" id="st-d0_s13" data-properties="anther fixation;anther architecture or structure in adjective form;anther dehiscence;slit dehiscence or orientation">anthers dorsifixed, 2-locular, extrorsely dehiscent by longitudinal slits.</span> <span class="statement" id="st-d0_s14" data-properties="flower architecture;bract architecture or arrangement or growth form;bracteole presence;bracteole presence;bracteole atypical quantity;bracteole quantity"><b>Pistillate </b>flowers subtended by solitary bract, bracteoles present or absent, usually 2-4 (-8);</span> <span class="statement" id="st-d0_s15" data-properties="pistil quantity;pistil architecture;pistil architecture or structure in adjective form">pistils 1, 2-carpellate, 1-locular;</span> <span class="statement" id="st-d0_s16" data-properties="ovule quantity;ovule position;ovule orientation">ovules 1, basal, erect;</span> <span class="statement" id="st-d0_s17" data-properties="style presence;style height or length or size">styles, if present, short;</span> <span class="statement" id="st-d0_s18" data-properties="stigma quantity">stigmas 2.</span> <span class="statement" id="st-d0_s19" data-properties="fruit architecture;fruit architecture;fruit architecture or pubescence or relief;fruit architecture or pubescence or relief"><b>Fruits </b>drupaceous or nutlike, smooth or often covered with warty protuberances, these commonly with waxy coating;</span> <span class="statement" id="st-d0_s20" data-properties="beak duration;beak size;bracteole duration;bracteole size">fruits sometimes enclosed by persistent, accrescent bracts and bracteoles.</span> <span class="statement" id="st-d0_s21" data-properties="endosperm quantity"><b>Seeds </b>with little or no endosperm;</span> <span class="statement" id="st-d0_s22" data-properties="embryo course;cotyledon quantity;cotyledon shape">embryo straight, with 2 planoconvex cotyledons.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Shrubs </b>or small trees, evergreen or deciduous, usually aromatic and resinous. <b>Roots</b> commonly with nitrogen-fixing nodules. <b>Leaves</b> alternate, simple or pinnatifid; stipules absent or present; petiole present. <b>Leaf</b> blade commonly with peltate, multicellular, glandular trichomes. <b>Inflorescences</b> axillary catkins; bracts present. <b>Flowers</b> usually unisexual, occasionally bisexual, staminate and pistillate flowers usually on different plants, occasionally on same plants; perianth absent. <b>Staminate</b> flowers subtended by solitary bract; stamens 2-14(-22), hypogynous or ± epigynous; filaments filiform, distinct or basally connate; anthers dorsifixed, 2-locular, extrorsely dehiscent by longitudinal slits. <b>Pistillate</b> flowers subtended by solitary bract, bracteoles present or absent, usually 2-4(-8); pistils 1, 2-carpellate, 1-locular; ovules 1, basal, erect; styles, if present, short; stigmas 2. <b>Fruits</b> drupaceous or nutlike, smooth or often covered with warty protuberances, these commonly with waxy coating; fruits sometimes enclosed by persistent, accrescent bracts and bracteoles. <b>Seeds</b> with little or no endosperm; embryo straight, with 2 plano-convex cotyledons.</span><!--
  
 
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|distribution=Widespread in temperate and subtropical regions
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|distribution=Widespread in temperate and subtropical regions.
 
|discussion=<p>Genera 2-4, species ca. 50 (2 genera, 8 species in the flora).</p><!--
 
|discussion=<p>Genera 2-4, species ca. 50 (2 genera, 8 species in the flora).</p><!--
--><p>Significant disagreement exists concerning the number of genera to be recognized in Myricaceae. Myrica in the broad sense is sometimes divided into three genera. Comptonia L'Héritier ex Aiton is often segregated on the basis of leaf type, presence of stipules, and the burlike fruits with 6-8 accrescent bracts and bracteoles. Morella Loureiro sometimes is elevated from its usual rank of subgenus to emphasize differences concerning position of the catkins, size of the staminate bracts, and appearance of the fruits (A. Chevalier 1901; J. R. Baird 1968). The real question is the appropriate rank at which recognition should be made (T. S. Elias 1971). I follow a traditional approach in recognizing just Myrica and Comptonia in North America.</p>
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--><p>Significant disagreement exists concerning the number of genera to be recognized in Myricaceae. <i>Myrica</i> in the broad sense is sometimes divided into three genera. <i>Comptonia</i> L'Héritier ex Aiton is often segregated on the basis of leaf type, presence of stipules, and the burlike fruits with 6-8 accrescent bracts and bracteoles. Morella Loureiro sometimes is elevated from its usual rank of subgenus to emphasize differences concerning position of the catkins, size of the staminate bracts, and appearance of the fruits (A. Chevalier 1901; J. R. Baird 1968). The real question is the appropriate rank at which recognition should be made (T. S. Elias 1971). I follow a traditional approach in recognizing just <i>Myrica</i> and <i>Comptonia</i> in North America.</p>
 
|tables=
 
|tables=
 
|references={{Treatment/Reference
 
|references={{Treatment/Reference
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|family=Myricaceae
 
|family=Myricaceae
 
|illustrator=John Myers
 
|illustrator=John Myers
|distribution=Widespread in temperate and subtropical regions
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|illustration copyright=Flora of North America Association
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|distribution=Widespread in temperate and subtropical regions.
 
|reference=baird1968a;chevalier1901a;elias1971a;sheffy1972a;wilbur1994a;youngken1919a
 
|reference=baird1968a;chevalier1901a;elias1971a;sheffy1972a;wilbur1994a;youngken1919a
 
|publication title=
 
|publication title=
 
|publication year=
 
|publication year=
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_100.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V3/V3_100.xml
|anther architecture or structure in adjective form=2-locular
 
|anther dehiscence=dehiscent
 
|anther fixation=dorsifixed
 
|beak duration=persistent
 
|beak presence=absent
 
|beak size=accrescent
 
|bract architecture or arrangement or growth form=solitary;solitary
 
|bracteole atypical quantity=4;8
 
|bracteole duration=persistent
 
|bracteole presence=absent;absent
 
|bracteole quantity=2;4
 
|bracteole size=accrescent
 
|cotyledon quantity=2
 
|cotyledon shape=planoconvex
 
|embryo course=straight
 
|endosperm quantity=no
 
|filament fusion=connate;distinct
 
|filament shape=filiform
 
|flower architecture=pistillate;staminate;pistillate;staminate
 
|flower reproduction=bisexual;unisexual
 
|fruit architecture=nutlike;drupaceous
 
|fruit architecture or pubescence or relief=often covered with warty protuberances;smooth
 
|glandular trichome architecture=multicellular;peltate
 
|leaf architecture=simple
 
|leaf arrangement=alternate
 
|leaf shape=pinnatifid
 
|ovule orientation=erect
 
|ovule position=basal
 
|ovule quantity=1
 
|perianth presence=absent
 
|petiole presence=absent
 
|pistil architecture=2-carpellate
 
|pistil architecture or structure in adjective form=1-locular
 
|pistil quantity=1
 
|shrub coating=resinous
 
|shrub duration=deciduous;evergreen
 
|shrub odor=aromatic
 
|slit dehiscence or orientation=longitudinal
 
|stamen atypical quantity=14;22
 
|stamen position=epigynous;hypogynous
 
|stamen quantity=2;14
 
|stigma quantity=2
 
|stipule presence=absent;absent
 
|style height or length or size=short
 
|style presence=absent
 
|tree coating=resinous
 
|tree duration=deciduous;evergreen
 
|tree odor=aromatic
 
|tree size=small
 
 
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Latest revision as of 22:45, 5 November 2020

Shrubs or small trees, evergreen or deciduous, usually aromatic and resinous. Roots commonly with nitrogen-fixing nodules. Leaves alternate, simple or pinnatifid; stipules absent or present; petiole present. Leaf blade commonly with peltate, multicellular, glandular trichomes. Inflorescences axillary catkins; bracts present. Flowers usually unisexual, occasionally bisexual, staminate and pistillate flowers usually on different plants, occasionally on same plants; perianth absent. Staminate flowers subtended by solitary bract; stamens 2-14(-22), hypogynous or ± epigynous; filaments filiform, distinct or basally connate; anthers dorsifixed, 2-locular, extrorsely dehiscent by longitudinal slits. Pistillate flowers subtended by solitary bract, bracteoles present or absent, usually 2-4(-8); pistils 1, 2-carpellate, 1-locular; ovules 1, basal, erect; styles, if present, short; stigmas 2. Fruits drupaceous or nutlike, smooth or often covered with warty protuberances, these commonly with waxy coating; fruits sometimes enclosed by persistent, accrescent bracts and bracteoles. Seeds with little or no endosperm; embryo straight, with 2 plano-convex cotyledons.

Distribution

Widespread in temperate and subtropical regions.

Discussion

Genera 2-4, species ca. 50 (2 genera, 8 species in the flora).

Significant disagreement exists concerning the number of genera to be recognized in Myricaceae. Myrica in the broad sense is sometimes divided into three genera. Comptonia L'Héritier ex Aiton is often segregated on the basis of leaf type, presence of stipules, and the burlike fruits with 6-8 accrescent bracts and bracteoles. Morella Loureiro sometimes is elevated from its usual rank of subgenus to emphasize differences concerning position of the catkins, size of the staminate bracts, and appearance of the fruits (A. Chevalier 1901; J. R. Baird 1968). The real question is the appropriate rank at which recognition should be made (T. S. Elias 1971). I follow a traditional approach in recognizing just Myrica and Comptonia in North America.

Lower Taxa

Key

1 Leaves entire or serrate-denticulate, stipules absent; fruiting catkins short-cylindric; bracteoles 2-6, broadly ovate and equal to or shorter than fruits, or absent. Myrica
1 Leaves pinnatifid, stipules present; fruiting catkins globose-ovoid; bracteoles 2 at anthesis, linear-subulate, accrescent, developing 4-8 tertiary bracteoles, these much exceeding fruit. Comptonia