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Eleocharis decumbens Eleocharis diandra Eleocharis elongata Eleocharis engelmannii Eleocharis equisetoides Eleocharis erythropoda Eleocharis fallax Eleocharis

depressipes, Quercus douglasii, Quercus dumosa, Quercus durata, Quercus engelmannii, Quercus fusiformis, Quercus gambelii, Quercus garryana, Quercus geminata

boomii, Isoëtes butleri, Isoëtes caroliniana, Isoëtes echinospora, Isoëtes engelmannii, Isoëtes flaccida, Isoëtes georgiana, Isoëtes howellii, Isoëtes lacustris

Polygonum californicum, Polygonum cascadense, Polygonum douglasii, Polygonum engelmannii, Polygonum heterosepalum, Polygonum hickmanii, Polygonum majus, Polygonum

Oenothera arizonica, Oenothera avita, Oenothera deltoides, Oenothera engelmannii, Oenothera neomexicana, Oenothera nuttallii, Oenothera pallida (Spach)

familyPinaceae genusPicea Show Lower Taxa Picea abies, Picea breweriana, Picea engelmannii, Picea glauca, Picea mariana, Picea pungens, Picea rubens, Picea sitchensis

coccineus, Echinocereus dasyacanthus, Echinocereus davisii, Echinocereus engelmannii, Echinocereus enneacanthus, Echinocereus fasciculatus, Echinocereus fendleri

Physaria dornii, Physaria douglasii, Physaria eburniflora, Physaria engelmannii, Physaria eriocarpa, Physaria fendleri, Physaria filiformis, Physaria

speciesOpuntia engelmannii Show Lower Taxa Opuntia engelmannii var. engelmannii, Opuntia engelmannii var. flavispina, Opuntia engelmannii var. flexispina

Synonyms: Eleocharis engelmannii var. detonsa A. Gray Eleocharis engelmannii var. monticola (Fernald) Svenson Eleocharis engelmannii var. robusta Fernald

LAYOUT:treatment:TUOYAL familyPinaceae genusPicea speciesPicea engelmannii Show Lower Taxa Picea engelmannii var. engelmannii Engelmann Trans. Acad. Sci. St. Louis 2: 212

Pinus coulteri, Pinus echinata, Pinus edulis, Pinus elliottii, Pinus engelmannii, Pinus flexilis, Pinus glabra, Pinus jeffreyi, Pinus lambertiana, Pinus

called I. engelmannii var. georgiana Engelmann. This variety may represent a tetraploid cytotype. A tetraploid population of I. engelmannii (2n = 44) from

Ovatae Show Lower Taxa Eleocharis aestuum, Eleocharis diandra, Eleocharis engelmannii, Eleocharis lanceolata, Eleocharis obtusa, Eleocharis ovata Svenson Rhodora

genusEucephalus Show Lower Taxa Eucephalus breweri, Eucephalus elegans, Eucephalus engelmannii, Eucephalus glabratus, Eucephalus glaucescens, Eucephalus gormanii, Eucephalus

chisosensis, Opuntia chlorotica, Opuntia cubensis, Opuntia ellisiana, Opuntia engelmannii, Opuntia ficus-indica, Opuntia fragilis, Opuntia humifusa, Opuntia littoralis

tribeAsteraceae tribe Astereae genusOönopsis Show Lower Taxa Oönopsis engelmannii, Oönopsis foliosa, Oönopsis multicaulis, Oönopsis wardii (Nuttall) Greene

Ophioglossum californicum, Ophioglossum crotalophoroides, Ophioglossum engelmannii, Ophioglossum nudicaule, Ophioglossum petiolatum, Ophioglossum pusillum

LAYOUT:treatment:TUOYAL familyOphioglossaceae genusOphioglossum speciesOphioglossum engelmannii Prantl Ber. Deutsch. Bot. Ges. 1: 351. 1883. Warren H. Wagner Jr., Florence

douglasii, Cirsium drummondii, Cirsium eatonii, Cirsium edule, Cirsium engelmannii, Cirsium flodmanii, Cirsium foliosum, Cirsium fontinale, Cirsium grahamii

familyHydrocharitaceae genusHalophila Show Lower Taxa Halophila decipiens, Halophila engelmannii, Halophila johnsonii Thouars Genera Nova Madagascariensia 2. 1806. Robert

elatior, Erigeron elatus, Erigeron elegantulus, Erigeron elmeri, Erigeron engelmannii, Erigeron evermannii, Erigeron eximius, Erigeron filifolius, Erigeron

genusCarex sectionCarex sect. Inflatae Show Lower Taxa Carex breweri, Carex engelmannii, Carex subnigricans Kükenthal in H. G. A. Engler, Pflanzenr. 20[IV,38]:

Daniel F. Brunton Common names: Carolina quillwort Basionym: Isoëtes engelmannii var. caroliniana A. A. Eaton Fern Bull 8: 60. 1900 Treatment appears in

Geraldine A. Allen Common names: Cascade aster Endemic Basionym: Aster engelmannii var. ledophyllus A. Gray Proc. Amer. Acad. Arts 8: 388. 1872 Synonyms:

Polygonoideae genusPolygonum sectionPolygonum sect. Duravia speciesPolygonum engelmannii Greene Bull. Calif. Acad. Sci. 1: 126. 1885. Mihai Costea, François J

Ophioglossaceae Ophioglossaceae Ophioglossaceae Ophioglossum vulgatum Ophioglossum engelmannii Ophioglossum pusillum Ophioglossum crotalophoroides Cheiroglossa palmata

Crataegus berberifolia var. berberifolia, Crataegus berberifolia var. engelmannii Torrey & A. Gray Fl. N. Amer. 1: 469. 1840. James B. Phipps Common names:

mm high in E. engelmannii var. robusta); perianth bristles often absent, rudimentary to slightly exceeding tubercle. Eleocharis engelmannii 23 Tubercles

represents an allotetraploid with I. engelmannii as one of its parents. Isoëtes louisianensis may represent I. engelmannii x melanopoda (B. M. Boom 1982). Spores

genusPicea speciesPicea engelmannii varietyPicea engelmannii var. engelmannii Engelmann Ronald J. Taylor Synonyms: Picea glauca subsp. engelmannii (Engelmann) T

E. engelmannii var. engelmannii. Similar plants from the opposite, eastern, side of the Sonoran Desert, in Arizona, have been called E. engelmannii var

Basionym: Bigelowia engelmannii A. Gray Proc. Amer. Acad. Arts 11: 75. 1876 (as Bigelovia engelmanni) Synonyms: Haplopappus engelmannii (A. Gray) H. M. Hall

varietyCrataegus berberifolia var. engelmannii (Sargent) Eggleston Rhodora 10: 76. 1908. James B. Phipps Endemic Basionym: Crataegus engelmannii Sargent Bot. Gaz. 31:

Aster elegans var. engelmannii D. C. Eaton in S. Watson, Botany (Fortieth Parallel), 144. 1871 (as engelmanni) Synonyms: Aster engelmannii (D. C. Eaton) A

Vesicaria engelmannii A. Gray Gen. Amer. Bor. 1: 162, plate 70. 1848 Synonyms: Alyssum engelmannii (A. Gray) Kuntze Lesquerella engelmannii (A. Gray) S

, Wyo. Carex engelmannii occurs north and east of the range of C. breweri; both species occur on Mt. Adams, Washington. Carex engelmannii is most similar

LAYOUT:treatment:TUOYAL familyHydrocharitaceae genusHalophila speciesHalophila engelmannii Ascherson in G. B. von Neumayer, Anleitung zu Wissenschaftlichen Beobachtungen

California). Quercus engelmannii is closely related to and possibly conspecific with Q. oblongifolia. The cups of Q. engelmannii are larger, deeper, and

Synonyms: Aloe yuccaefolia A. Gray Hesperaloe engelmannii Krauskopf Hesperaloe parviflora var. engelmannii (Krauskopf) Trelease Hesperaloe yuccaefolia Treatment

Brown, Ill. Fl. N. U.S. 2: 137, fig. 1749. 1897 Synonyms: Lesquerella engelmannii subsp. ovalifolia (Rydberg ex Britton) C. Clark Treatment appears in FNA

Synonyms: Selaginella densa var. scopulorum (Maxon) R. M. Tryon Selaginella engelmannii var. scopulorum (Maxon) C. F. Reed Treatment appears in FNA Volume 2.

Y., S.Dak., Vt., Wis., Wyo. In areas of sympatry Picea glauca and P. engelmannii regularly hybridize and intergrade completely (R.Daubenmire 1974; E.H

Myers Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Picea breweriana Picea engelmannii Picea rubens Picea glauca Picea sitchensis John Myers John Myers John

elevational limits of this variety, occasional intermediates with Q. engelmannii occur [= Q. ×grandidentata Ewan (as species)]. These putative hybrids

subfam. Opuntioideae genusOpuntia speciesOpuntia engelmannii varietyOpuntia engelmannii var. engelmannii Donald J. Pinkava Common names: Engelmann’s pricklypear

subfamilyCactaceae subfam. Opuntioideae genusOpuntia speciesOpuntia engelmannii varietyOpuntia engelmannii var. lindheimeri (Engelmann) B. D. Parfitt & Pinkava Madroño

subfamilyCactaceae subfam. Opuntioideae genusOpuntia speciesOpuntia engelmannii varietyOpuntia engelmannii var. linguiformis (Griffiths) B. D. Parfitt & Pinkava Madroño

subfamilyCactaceae subfam. Opuntioideae genusOpuntia speciesOpuntia engelmannii varietyOpuntia engelmannii var. flexispina (Griffiths) B. D. Parfitt & Pinkava Madroño

subfamilyCactaceae subfam. Opuntioideae genusOpuntia speciesOpuntia engelmannii varietyOpuntia engelmannii var. flavispina (L. D. Benson) B. D. Parfitt & Pinkava Madroño

collection from this locality was reported by B. M. Boom (1982) to be I. engelmannii x piedmontana, but the broad velum and bold texture of the megaspores

primarily Pinus ponderosa, sometimes P. arizonica, P. durangensis, and P. engelmannii); w United States. Arceuthobium vaginatum

speciesOenothera engelmannii (Small) Munz Amer. J. Bot. 18: 316. 1931. (as engelmanni) Warren L. Wagner Basionym: Anogra pallida var. engelmannii Small Bull

Ferguson) Croizat C. capitatus var. albinoides (A. M. Ferguson) Shinners C. engelmannii var. albinoides A. M. Ferguson C. muelleri var. albinoides (A. M. Ferguson)

compact habit of Erigeron engelmannii but sparsely spreading-hirsute stems; they may be intermediate between E. engelmannii and E. concinnus. Unusual variation

Watson Smithsonian Misc. Collect. 258: 118. 1878 Synonyms: Echinocereus engelmannii subsp. fasciculatus (Engelmann ex S. Watson) Blume & Mich. Lange Echinocereus

A. Allen Common names: Olympic Mountain aster Endemic Basionym: Aster engelmannii var. paucicapitatus B. L. Robinson Proc. Amer. Acad. Arts 26: 176. 1891

Common names: Klickitat aster EndemicConservation concern Basionym: Aster engelmannii var. glaucescens A. Gray in A. Gray et al., Syn. Fl. N. Amer. 1(2): 200

LAYOUT:treatment:TUOYAL familyPinaceae genusPinus speciesPinus engelmannii Carrière 3: 227. 1854. Robert Kral Common names: Apache pine Basionym: Pinus

familyAsteraceae tribeAsteraceae tribe Cardueae genusCirsium speciesCirsium engelmannii Rydberg Fl. Rocky Mts., 1069. 1917. David J. Keil Common names: Blackland

WeedyIllustratedEndemic Synonyms: Acetosa hastatula (Baldwin) Á. Löve Rumex engelmannii Meisner Treatment appears in FNA Volume 5. Treatment on page 502. Mentioned

Coryphantha scheeri var. valida (Schott ex Engelmann) L. D. Benson Mammillaria engelmannii Treatment appears in FNA Volume 4. Mentioned on page 226. Plants unbranched

Basionym: Erigeron engelmannii subsp. davisii Cronquist Leafl. W. Bot. 3: 167. 1942 (as engelmanni) Synonyms: Erigeron engelmannii var. davisii (Cronquist)

gracilescens (Rechinger f.) Á. Löve Acetosella paucifolia (Nuttall) Á. Löve Rumex engelmannii var. geyeri Meisner Rumex geyeri (Meisner) Trelease Rumex paucifolius

John K. Morton Common names: Bridges’s catchfly Endemic Synonyms: Silene engelmannii Rohrbach Silene incompta A. Gray Silene longistylis Engelmann ex S. Watson

stiff leaves. Isoëtes tuckermanii hybridizes with I. echinospora; I. engelmannii [ = I. x foveolata A. A. Eaton ex Dodge]; and I. lacustris [ = I. x harveyi

ridges on the distal hemispheres. Isoëtes flaccida hybridizes with I. engelmannii None. None. window.propertiesFromHigherTaxa=[{"rank":"genus","name":"Iso\u00ebtes"

riparia hybridizes with I. echinospora [ = I. x dodgei A. A. Eaton]; I. engelmannii [ = I. x brittonii Brunton & W. C. Taylor]; I. lacustris [ = I. x jeffreyi

northeastern North America. Isoëtes echinospora hybridizes with I. bolanderi; I. engelmannii [ = I. x eatonii Dodge (later synonym = I. x gravesii A. A. Eaton)]; I

hibridizes with I. echinospora [ = I. x hickeyi W. C. Taylor & Luebke]; I. engelmannii; I. riparia [ = I. x jeffreyi D. M. Britton & Brunton]; and with I. tuckermanii

Watson Cyperus eggersii Boeckeler Cyperus engelmannii Steudel Cyperus ferax Richard Cyperus ferax subsp. engelmannii (Steudel) Kükenthal Cyperus ferax subsp

Proc. Amer. Acad. Arts 3: 291. 1856. Donald J. Pinkava Synonyms: Opuntia engelmannii var. occidentalis (Engelmann) Engelmann Treatment appears in FNA Volume

Nichol’s hedgehog cactus golden hedgehog cactus Basionym: Echinocereus engelmannii var. nicholii L. D. Benson Proc. Calif. Acad. Sci., ser. 4, 25: 258, plate

Steven E. Clemants* Endemic Synonyms: Juncus echinatus Muhlenberg Juncus engelmannii Buchenau Treatment appears in FNA Volume 22. Herbs, perennial, cespitose

ovalifolia var. alba Goodman Rhodora 38: 239. 1936 Synonyms: Lesquerella engelmannii subsp. alba (Goodman) C. Clark Lesquerella ovalifolia subsp. alba (Goodman)

Myers Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Picea breweriana Picea engelmannii Picea rubens Picea glauca Picea sitchensis John Myers John Myers John

Acad. Sci. 4: 15. 1905. Donald J. Pinkava Illustrated Basionym: Opuntia engelmannii var. littoralis Engelmann in W. H. Brewer et al., Bot. Calif. 1: 248.

Ophioglossaceae Ophioglossaceae Ophioglossaceae Ophioglossum vulgatum Ophioglossum engelmannii Ophioglossum pusillum Ophioglossum crotalophoroides Cheiroglossa palmata

with E. ovata and E. aestuum. A few specimens are intermediate with E. engelmannii. Eleocharis obtusa is sometimes treated as conspecific with E. ovata,

laricis; secondary to rare hosts include Abies grandis, A. lasiocarpa, Picea engelmannii, Pinus albicaulis, P. contorta, P. monticola, and P. ponderosa. This dwarf

Subspecies microcarpum is a serious pathogen on its principal hosts, Picea engelmannii and P. pungens, as well as on Pinus aristata in northern Arizona. Rare

also on Abies amabilis, A. concolor, A. grandis, A. lasiocarpa, Picea engelmannii, and P. pungens. Plants of Arceuthobium douglasii induce massive systemic

incidence of infection. Additional secondary to rare hosts include Picea engelmannii, Pinus balfouriana, P. contorta, P. monticola, P. ponderosa, and Tsuga

Secondary to rare hosts include Abies grandis, Picea breweriana, Picea engelmannii, Picea sitchensis, Pinus albicaulis, Pinus monticola, and Pseudotsuga

dry habitat preference; putative hybrids are known, however, with Q. engelmannii and Q. lobata. Some populations of Q. berberidifolia from higher elevations

parasitizes other principal hosts such as P. arizonica, P. durangensis, and P. engelmannii as well as secondary and occasional hosts such as P. aristata, P. contorta

Utah, Wyo. Limited hybridization occurs between Picea pungens and P. engelmannii (R.Daubenmire 1972; R.J. Taylor et al. 1975). Blue spruce (Picea pungens)

north to Mt. Hood. A. Cronquist (1969) considered Carex breweri and C. engelmannii conspecific; he distinguished them at the varietal level by a difference

subsp. melina and R. acicularis subsp. sayi has been recognized as R. ×engelmannii S. Watson. It is based on such intermediate characteristics as few infrastipular

assigned to varieties of Opuntia phaeacantha has been segregated as O. engelmannii. Various combinations of overlapping character states make separation

tuberculate cups. Of greater difficulty are swarms of putative hybrids with Q. engelmannii, sometimes referred to as Q. ×acutidens. In Joshua Tree National Monument

Service 1983), which has characteristics of both Picea glauca and P. engelmannii. These trees can similarly be called interior subalpine fir, i.e., A.

durata var. gabrielensis, Quercus ellipsoidalis, Quercus emoryi, Quercus engelmannii, Quercus falcata, Quercus fusiformis, Quercus gambelii, Quercus garryana

apparently also occur in the same area. Some plants seem to approach E. engelmannii in their reduced vestiture. Study is needed to evaluate the evolutionary

Mexico (Chihuahua). Opuntia macrorhiza hybridizes with the hexaploid O. engelmannii (apparently var. lindheimeri) forming the pentaploid O. ×edwardsii V.

(introduced), Australia (introduced). Opuntia stricta hybridizes with O. engelmannii (apparently var. lindheimeri) forming O. ×alta Griffiths (as species)

similarity to some of the tree forms that are putative hybrids between Q. engelmannii and Q. cornelius-mulleri in San Diego County. The latter populations,

important element there, in particular Pinus cooperi, P. durangensis, P. engelmannii, P. leiophylla, P. lumholtzii, and Pinus ponderosa var. arizonica (D.E

parkland with trees of lodgepole pine and probably Engelmann spruce (Picea engelmannii) (Davis Lake and Battle Ground Lake, fig. 4.13; C.W. Barnosky et al. 1987)

turn in the uppermost forests is replaced by Abies bifolia and Picea engelmannii. Especially at middle elevations, Populus tremuloides and the fire-dependent