Difference between revisions of "Thelypteridaceae"

Ching ex Pichi Sermolli
Common names: Marsh Fern Family
Treatment appears in FNA Volume 2. Treatment on page 206.
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--><span class="statement" id="st-d0_s0" data-properties="plant growth form or habitat;plant growth form or habitat"><b>Plants </b>terrestrial or on rock [epiphytic].</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stem orientation;stem architecture or pubescence"><b>Stems </b>creeping to erect, scaly at apex.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf growth form"><b>Leaves </b>monomorphic or somewhat dimorphic [dimorphic].</span> <span class="statement" id="st-d0_s3" data-properties="vascular-bundle count;vascular-bundle shape"><b>Petiole </b>in cross-section with 2 crescent-shaped vascular-bundles at base.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade shape;blade shape;blade architecture or shape"><b>Blade </b>pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple];</span> <span class="statement" id="st-d0_s5" data-properties="rachis architecture;groove architecture">rachis grooved adaxially or not, grooves not continuous with grooves of next order.</span> <span class="statement" id="st-d0_s6" data-properties="vein fusion;vein architecture;veinlet fusion"><b>Veins </b>free or anastomosing, running to margin, areoles with or without included free veinlets.</span> <span class="statement" id="st-d0_s7" data-properties="hair coloration;hair architecture or shape;hair shape;hair architecture;hair arrangement or shape;hair count"><b>Indument </b>of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking.</span> <span class="statement" id="st-d0_s8" data-properties="sorus position;sorus position;sorus position;sorus position;sorus shape;sorus shape;sorus shape"><b>Sori </b>inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins;</span> <span class="statement" id="st-d0_s9" data-properties="indusium shape;indusium count">indusia reniform or sometimes absent.</span> <span class="statement" id="st-d0_s10" data-properties="spore arrangement;spore architecture;perispore prominence;perispore shape;perispore architecture;perispore architecture"><b>Spores </b>bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore.</span> <span class="statement" id="st-d0_s11" data-properties="gametophyte coloration;gametophyte shape;gametophyte pubescence;gametophyte pubescence"><b>Gametophytes </b>green, cordate, usually hairy or glandular;</span> <span class="statement" id="st-d0_s12" data-properties="antheridium architecture">antheridia 3-celled.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>terrestrial or on rock [epiphytic]. <b>Stems</b> creeping to erect, scaly at apex. <b>Leaves</b> monomorphic or somewhat dimorphic [dimorphic]. <b>Petiole</b> in cross section with 2 crescent-shaped vascular bundles at base. <b>Blade</b> pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. <b>Veins</b> free or anastomosing, running to margin, areoles with or without included free veinlets. <b>Indument</b> of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. <b>Sori</b> inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. <b>Spores</b> bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. <b>Gametophytes</b> green, cordate, usually hairy or glandular; antheridia 3-celled.</span><!--
  
 
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_797.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_797.xml
|antheridium architecture=3-celled
 
|blade architecture or shape=2+-pinnate
 
|blade shape=pinnate;pinnate-pinnatifid
 
|gametophyte coloration=green
 
|gametophyte pubescence=glandular;hairy
 
|gametophyte shape=cordate
 
|groove architecture=not continuous
 
|hair architecture=septate
 
|hair architecture or shape=needlelike
 
|hair arrangement or shape=stellate
 
|hair coloration=transparent
 
|hair count=lacking
 
|hair shape=hooked
 
|indusium count=absent
 
|indusium shape=reniform
 
|leaf architecture=monomorphic
 
|leaf growth form=dimorphic
 
|perispore architecture=reticulate;echinate
 
|perispore prominence=prominent
 
|perispore shape=crested
 
|plant growth form or habitat=on rock;terrestrial
 
|rachis architecture=grooved
 
|sorus position=marginal;inframedial;supramedial
 
|sorus shape=elongate;oblong;round
 
|spore architecture=monolete
 
|spore arrangement=bilateral
 
|stem architecture or pubescence=scaly
 
|stem orientation=creeping;erect
 
|vascular-bundle count=2
 
|vascular-bundle shape=crescent-shaped
 
|vein architecture=anastomosing
 
|vein fusion=free
 
|veinlet fusion=free
 
 
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Revision as of 13:28, 27 July 2019

Plants terrestrial or on rock [epiphytic]. Stems creeping to erect, scaly at apex. Leaves monomorphic or somewhat dimorphic [dimorphic]. Petiole in cross section with 2 crescent-shaped vascular bundles at base. Blade pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. Veins free or anastomosing, running to margin, areoles with or without included free veinlets. Indument of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. Sori inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. Spores bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. Gametophytes green, cordate, usually hairy or glandular; antheridia 3-celled.

Discussion

Members of Thelypteridaceae have historically been associated with Dryopteridaceae (in particular, Dryopteris) but in fact have no close relationship with that family. Thelypteris and allies differ from Dryopteris and allies by their indument of transparent needlelike hairs (versus needlelike hairs absent in Dryopteridaceae); general absence of blade scales (versus blade scales often present); petiole vasculature in cross section with two crescent-shaped bundles (versus many round bundles arranged in an arc, Athyrium and allies exceptional); generally 1-pinnate to pinnate-pinnatifid blades (versus often more divided); veins usually not forking in the ultimate segments (versus often forking); adaxial grooves discontinuous from rachis to costae, or grooves lacking (versus grooves often continuous); and chromosome base numbers from 27–36 (versus generally 40, 41).

Genera 1 to ca. 30, depending on circumscription, species ca. 900 (as circumscribed here, 3 genera and 25 species in the flora).

Selected References

None.

Key

1 Blades 1-pinnate to deeply pinnate-pinnatifid; costae grooved adaxially; veins meeting margin at or above sinus or united below sinus. Thelypteris
1 Blades 2-pinnatifid, with pinnae at least in distal 1/2 of blade connected by wings along rachis, or blades 2-pinnate-pinnatifid; costae not grooved adaxially; veins commonly meeting margin above sinus. > 2
2 Pinnae free, rachis not winged; blades 2-pinnate or more divided; costal hairs septate, often longer than 1 mm; indusia small, less than 0.3 mm diam. Macrothelypteris
2 Pinnae mostly connected by wings along rachis, the wings often forming semicircular lobes between pinnae; blades 2-pinnatifid; costal hairs not septate, shorter than 0.5 mm; indusia absent. Phegopteris