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Paysonia
lyrata, Paysonia perforata, Paysonia stonensis O’Kane & Al-Shehbaz Novon 12: 380. 2002. Steve L. O’Kane Jr. Etymology: For Edwin Blake Payson, 1893–1927

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Physaria
(Nuttall ex Torrey & A. Gray) A. Gray Gen. Amer. Bor. 1: 162. 1848. Steve L. O’Kane Jr. Common names: Bladderpod Etymology: Greek physa, bladder, alluding

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Physaria kingii
Physaria kingii subsp. utahensis (S. Watson) O’Kane & Al-Shehbaz Novon 12: 324. 2002. Steve L. O’Kane Jr. Common names: King bladderpod Basionym: Vesicaria

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Physaria argyraea
Taxa Physaria argyraea subsp. argyraea (A. Gray) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Silver bladderpod Basionym: Vesicaria

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Physaria carinata
Physaria carinata subsp. pulchella (Rollins) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Idaho bladderpod Basionym: Lesquerella

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Physaria douglasii
Physaria douglasii subsp. tuplashensis (S. Watson) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Douglas’s bladderpod Basionym:

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Physaria hitchcockii
Physaria hitchcockii subsp. rubicundula (Munz) O’Kane & Al-Shehbaz Novon 12: 324. 2002. Steve L. O’Kane Jr. Common names: Hitchcock’s bladderpod Basionym:

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Physaria arenosa
Physaria arenosa subsp. argillosa (Richardson) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Great Plains bladderpod Basionym:

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Physaria didymocarpa
didymocarpa subsp. lyrata (Hooker) A. Gray Gen. Amer. Bor. 1: 162. 1848. Steve L. O’Kane Jr. Common names: Common twinpod Endemic Basionym: Vesicaria didymocarpa Hooker

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Physaria hemiphysaria
Physaria hemiphysaria subsp. lucens (Maguire) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Intermountain or skyline bladderpod

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Physaria occidentalis
occidentalis subsp. occidentalis (S. Watson) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Common names: Western bladderpod Basionym: Vesicaria

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Physaria gracilis
Physaria gracilis subsp. nuttallii (Hooker) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Spreading bladderpod Basionym:

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Paysonia lasiocarpa
lasiocarpa subsp. lasiocarpa (Hooker ex A. Gray) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Roughpod bladderpod Basionym: Vesicaria

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Physaria saximontana
saximontana subsp. saximontana Rollins Contr. Gray Herb. 214: 13. 1984. Steve L. O’Kane Jr. Common names: Rocky Mountain or Fremont County twinpod Endemic Treatment

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Physaria floribunda
floribunda subsp. osterhoutii Rydberg Bull. Torrey Bot. Club 28: 279. 1901. Steve L. O’Kane Jr. Common names: Point-tip twinpod Treatment appears in FNA Volume 7

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Physaria geyeri
geyeri subsp. purpurea (Hooker) A. Gray Gen. Amer. Bor. 1: 162. 1848. Steve L. O’Kane Jr. Common names: Geyer’s bladderpod IllustratedEndemic Basionym: Vesicaria

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Physaria lepidota
membranacea Rollins Brittonia 33: 335, figs. 1, 2. 1981. Steve L. O’Kane Jr. Common names: Kane County twinpod Endemic Treatment appears in FNA Volume 7

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Physaria ovalifolia
(Rydberg) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Common names: Roundleaf bladderpod Basionym: Lesquerella ovalifolia Rydberg in N. L. Britton

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Physaria douglasii subsp. douglasii
speciesPhysaria douglasii subspeciesPhysaria douglasii subsp. douglasii Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 635. Cauline

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Physaria newberryi
subsp. yesicola A. Gray in J. C. Ives, Rep. Colorado R. 4: 6. 1861. Steve L. O’Kane Jr. Common names: Newberry twinpod Synonyms: Physaria didymocarpa var

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Physaria carinata subsp. pulchella
subspeciesPhysaria carinata subsp. pulchella (Rollins) O’Kane Novon 17: 379. 2007. Steve L. O’Kane Jr. Common names: Beautiful bladderpod Basionym: Lesquerella

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Physaria carinata subsp. paysonii
subspeciesPhysaria carinata subsp. paysonii (Rollins) O’Kane Novon 17: 379. 2007. Steve L. O’Kane Jr. Common names: Payson’s bladderpod Basionym: Lesquerella

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Physaria didymocarpa subsp. lyrata
subspeciesPhysaria didymocarpa subsp. lyrata (C. L. Hitchcock) O’Kane Novon 17: 379. 2007. Steve L. O’Kane Jr. Common names: Idaho or salmon twinpod Basionym:

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Physaria arenosa subsp. argillosa
arenosa subsp. argillosa (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Secund or sidesaddle bladderpod

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Physaria didymocarpa subsp. lanata
subspeciesPhysaria didymocarpa subsp. lanata (A. Nelson) O’Kane Novon 17: 379. 2007. Steve L. O’Kane Jr. Common names: Lanate common twinpod Basionym: Physaria

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Physaria didymocarpa subsp. didymocarpa
speciesPhysaria didymocarpa subspeciesPhysaria didymocarpa subsp. didymocarpa Steve L. O’Kane Jr. Synonyms: Physaria macrantha Blankinship Treatment appears in FNA

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Paysonia lasiocarpa subsp. lasiocarpa
speciesPaysonia lasiocarpa subspeciesPaysonia lasiocarpa subsp. lasiocarpa Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 614. Illustrator:

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Physaria carinata subsp. carinata
speciesPhysaria carinata subspeciesPhysaria carinata subsp. carinata Steve L. O’Kane Jr. Synonyms: Lesquerella carinata var. languida Rollins Treatment appears

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Physaria arenosa subsp. arenosa
speciesPhysaria arenosa subspeciesPhysaria arenosa subsp. arenosa Steve L. O’Kane Jr. Synonyms: Lesquerella lunellii A. Nelson Lesquerella macounii Greene

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Physaria douglasii subsp. tuplashensis
subspeciesPhysaria douglasii subsp. tuplashensis (Rollins Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: White Bluffs bladderpod Basionym: Lesquerella tuplashensis Rollins

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Physaria curvipes
genusPhysaria speciesPhysaria curvipes (A. Nelson) Grady & O’Kane Novon 17: 183. 2007. Steve L. O’Kane Jr. Common names: Curved bladderpod Basionym: Lesquerella

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Paysonia lasiocarpa subsp. berlandieri
subspeciesPaysonia lasiocarpa subsp. berlandieri (A. Gray) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Berlandier bladderpod Basionym:

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Physaria eriocarpa
genusPhysaria speciesPhysaria eriocarpa Grady & O’Kane Novon 17: 184, fig. 3. 2007. Steve L. O’Kane Jr. Common names: Sheep Mountain bladderpod Treatment

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Physaria cinerea
genusPhysaria speciesPhysaria cinerea (S. Watson) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Basin bladderpod Basionym: Lesquerella

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Physaria congesta
genusPhysaria speciesPhysaria congesta (Rollins) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Dudley Bluffs bladderpod Basionym:

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Physaria lesicae
genusPhysaria speciesPhysaria lesicae (Rollins) O’Kane & Al-Shehbaz Novon 12: 325. 2002 (as lesicii). Steve L. O’Kane Jr. Common names: Pryor Mountains bladderpod

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Paysonia auriculata
speciesPaysonia auriculata (Engelmann & A. Gray) O’Kane & Al-Shehbaz Novon 12: 380. 2002. Steve L. O’Kane Jr. Common names: Earleaf bladderpod Basionym: Vesicaria

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Physaria calcicola
genusPhysaria speciesPhysaria calcicola (Rollins) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Rocky Mountain bladderpod Basionym:

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Physaria densiflora
genusPhysaria speciesPhysaria densiflora (A. Gray) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Denseflower or low bladderpod Basionym:

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Paysonia densipila
genusPaysonia speciesPaysonia densipila (Rollins) O’Kane & Al-Shehbaz Novon 12: 380. 2002. Steve L. O’Kane Jr. Common names: Duck River bladderpod Basionym:

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Paysonia lyrata
genusPaysonia speciesPaysonia lyrata (Rollins) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Lyrate or lyreleaf bladderpod Basionym:

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Physaria angustifolia
speciesPhysaria angustifolia (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Threadleaf bladderpod Basionym:

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Physaria arctica
speciesPhysaria arctica (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Arctic bladderpod Basionym: Alyssum

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Physaria aurea
genusPhysaria speciesPhysaria aurea (Wooton) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Golden bladderpod Basionym: Lesquerella

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Physaria calderi
speciesPhysaria calderi (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Calder’s bladderpod Basionym: Lesquerella

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Physaria engelmannii
genusPhysaria speciesPhysaria engelmannii (A. Gray) O’Kane & Al-Shehbaz Novon 12: 322. 2002. Steve L. O’Kane Jr. Common names: Engelmann’s bladderpod Basionym:

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Paysonia grandiflora
genusPaysonia speciesPaysonia grandiflora (Hooker) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Bigflower bladderpod Basionym:

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Physaria argyraea subsp. argyraea
speciesPhysaria argyraea subspeciesPhysaria argyraea subsp. argyraea Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 627. Perennials;

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Physaria chambersii
556, figs. 15–18. 1939. Steve L. O’Kane Jr. Common names: Chambers’s twinpod Endemic Synonyms: Physaria chambersii var. canaani S. L. Welsh Physaria chambersii

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Physaria acutifolia
28: 279. 1901. Steve L. O’Kane Jr. Common names: Rydberg’s or sharpleaf twinpod Synonyms: Physaria acutifolia var. stylosa (Rollins) S. L. Welsh Physaria

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Physaria cordiformis
speciesPhysaria cordiformis Rollins Contr. Gray Herb. 171: 47. 1950. Steve L. O’Kane Jr. Common names: Wassuk Range bladderpod Synonyms: Lesquerella cordiformis (Rollins)

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Physaria condensata
condensata Rollins Rhodora 41: 407, plate 556, figs. 1, 9, 10. 1939. Steve L. O’Kane Jr. Common names: Tufted twinpod Treatment appears in FNA Volume 7. Treatment

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Physaria bellii
Mulligan Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. 1966. Steve L. O’Kane Jr. Common names: Bell’s or Front Range twinpod Treatment appears in

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Physaria eburniflora
genusPhysaria speciesPhysaria eburniflora Rollins Brittonia 33: 333. 1981. Steve L. O’Kane Jr. Common names: Devils Gate twinpod Endemic Treatment appears in FNA

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Physaria brassicoides
speciesPhysaria brassicoides Rydberg Bull. Torrey Bot. Club 29: 237. 1902. Steve L. O’Kane Jr. Common names: Double bladderpod IllustratedEndemic Treatment appears

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Physaria dornii
speciesPhysaria dornii Lichvar Brittonia 35: 150, figs. 1–3. 1983. Steve L. O’Kane Jr. Common names: Dorn’s twinpod EndemicConservation concern Treatment

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Physaria alpestris
genusPhysaria speciesPhysaria alpestris Suksdorf W. Amer. Sci. 15: 58. 1906. Steve L. O’Kane Jr. Common names: Washington or alpine twinpod Endemic Synonyms: Lesquerella

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Physaria geyeri subsp. geyeri
genusPhysaria speciesPhysaria geyeri subspeciesPhysaria geyeri subsp. geyeri Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 639. Trichomes

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Physaria saximontana subsp. saximontana
speciesPhysaria saximontana subspeciesPhysaria saximontana subsp. saximontana Steve L. O’Kane Jr. Common names: Fremont County bladderpod Treatment appears in FNA

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Physaria arizonica
genusPhysaria speciesPhysaria arizonica (S. Watson) O’Kane & Al-Shehbaz Novon 12: 321. 2002. Steve L. O’Kane Jr. Common names: Arizona bladderpod Basionym: Lesquerella

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Paysonia lescurii
genusPaysonia speciesPaysonia lescurii (A. Gray) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Lescur’s bladderpod Basionym: Vesicaria

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Paysonia stonensis
genusPaysonia speciesPaysonia stonensis (Rollins) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Stones River bladderpod Basionym:

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Paysonia perforata
genusPaysonia speciesPaysonia perforata (Rollins) O’Kane & Al-Shehbaz Novon 12: 381. 2002. Steve L. O’Kane Jr. Common names: Spring Creek bladderpod Basionym:

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Physaria alpina
genusPhysaria speciesPhysaria alpina Rollins Brittonia 33: 339. 1981. Steve L. O’Kane Jr. Common names: Avery Peak or alpine twinpod EndemicConservation concern

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Physaria hemiphysaria subsp. lucens
subspeciesPhysaria hemiphysaria subsp. lucens (S. L. Welsh & Reveal) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Tavaputs bladderpod Basionym:

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Physaria hitchcockii subsp. rubicundula
subspeciesPhysaria hitchcockii subsp. rubicundula (Rollins) O’Kane & Al-Shehbaz Novon 12: 324. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella rubicundula Rollins Contr

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Physaria lepidota subsp. membranacea
subspeciesPhysaria lepidota subsp. membranacea (Rollins) O’Kane Novon 17: 531. 2007. Steve L. O’Kane Jr. Basionym: Physaria chambersii var. membranacea Rollins

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Physaria newberryi subsp. yesicola
subspeciesPhysaria newberryi subsp. yesicola (Sivinski) O’Kane Novon 17: 381. 2007. Steve L. O’Kane Jr. Basionym: Physaria newberryi var. yesicola Sivinski

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Physaria saximontana subsp. dentata
subspeciesPhysaria saximontana subsp. dentata (Rollins) O’Kane Novon 17: 381. 2007. Steve L. O’Kane Jr. Basionym: Physaria saximontana var. dentata Rollins

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Physaria geyeri subsp. purpurea
subspeciesPhysaria geyeri subsp. purpurea (Rollins) O’Kane Novon 17: 380. 2007. Steve L. O’Kane Jr. Common names: Purple twinpod Basionym: Physaria geyeri

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Physaria floribunda subsp. osterhoutii
subspeciesPhysaria floribunda subsp. osterhoutii (Payson) O’Kane Novon 17: 379. 2007. Steve L. O’Kane Jr. Common names: Osterhout’s twinpod Basionym: Physaria

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Physaria hitchcockii subsp. confluens
hitchcockii subsp. confluens (Maguire & A. H. Holmgren) O’Kane Novon 17: 380. 2007. Steve L. O’Kane Jr. Basionym: Lesquerella hitchcockii subsp. confluens Maguire

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Physaria floribunda subsp. floribunda
speciesPhysaria floribunda subspeciesPhysaria floribunda subsp. floribunda Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 637. Racemes

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Physaria hemiphysaria subsp. hemiphysaria
speciesPhysaria hemiphysaria subspeciesPhysaria hemiphysaria subsp. hemiphysaria Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 642. Fruiting

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Physaria hitchcockii subsp. hitchcockii
speciesPhysaria hitchcockii subspeciesPhysaria hitchcockii subsp. hitchcockii Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 643. Plants

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Physaria lepidota subsp. lepidota
speciesPhysaria lepidota subspeciesPhysaria lepidota subsp. lepidota Steve L. O’Kane Jr. Treatment appears in FNA Volume 7. Treatment on page 648. Trichomes:

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Physaria gracilis subsp. gracilis
speciesPhysaria gracilis subspeciesPhysaria gracilis subsp. gracilis Steve L. O’Kane Jr. Synonyms: Lesquerella polyantha (Schlechtendal) Small Vesicaria

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Physaria newberryi subsp. newberryi
speciesPhysaria newberryi subspeciesPhysaria newberryi subsp. newberryi Steve L. O’Kane Jr. Synonyms: Physaria newberryi var. racemosa Rollins Treatment appears

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Physaria kingii subsp. latifolia
subspeciesPhysaria kingii subsp. latifolia (A. Nelson) O’Kane & Al-Shehbaz Novon 12: 325. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella latifolia A. Nelson Bot

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Physaria kingii subsp. utahensis
subspeciesPhysaria kingii subsp. utahensis (Rydberg) O’Kane Novon 17: 380. 2007. Steve L. O’Kane Jr. Common names: Utah bladderpod Basionym: Lesquerella

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Physaria kingii subsp. diversifolia
subspeciesPhysaria kingii subsp. diversifolia (Greene) O’Kane & Al-Shehbaz Novon 12: 325. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella diversifolia Greene Pittonia

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Physaria occidentalis subsp. cinerascens
subsp. cinerascens (Maguire & A. H. Holmgren) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella occidentalis var. cinerascens Maguire

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Physaria parvula
genusPhysaria speciesPhysaria parvula (Greene) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Common names: Pygmy bladderpod Basionym: Lesquerella

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Physaria kingii subsp. cobrensis
kingii subsp. cobrensis (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz Novon 12: 324. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella kingii var. cobrensis Rollins

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Physaria kingii subsp. bernardina
subspeciesPhysaria kingii subsp. bernardina (Munz) O’Kane & Al-Shehbaz Novon 12: 324. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella bernardina Munz Bull. S

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Physaria ovalifolia subsp. alba
subspeciesPhysaria ovalifolia subsp. alba (Goodman) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Basionym: Lesquerella ovalifolia var. alba Goodman

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Physaria gracilis subsp. nuttallii
gracilis subsp. nuttallii (Torrey & A. Gray) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Nuttall’s bladderpod Basionym:

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Physaria pycnantha
genusPhysaria speciesPhysaria pycnantha Grady & O’Kane Novon 17: 188, fig. 5. 2007. Steve L. O’Kane Jr. Common names: Mountain-view bladderpod Treatment

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Physaria thamnophila
speciesPhysaria thamnophila (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz Novon 12: 328. 2002. Steve L. O’Kane Jr. Common names: Zapata bladderpod Basionym: Lesquerella

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Physaria garrettii
genusPhysaria speciesPhysaria garrettii (Payson) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Garrett’s bladderpod Basionym:

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Physaria klausii
genusPhysaria speciesPhysaria klausii (Rollins) O’Kane & Al-Shehbaz Novon 12: 325. 2002. Steve L. O’Kane Jr. Common names: Rogers Pass or Klaus’s or Divide

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Physaria pachyphylla
Physarieae genusPhysaria speciesPhysaria pachyphylla O’Kane & Grady Novon 17: 187, fig. 4. 2007. Steve L. O’Kane Jr. Common names: Thick-leaf bladderpod Treatment

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Physaria pendula
genusPhysaria speciesPhysaria pendula (Rollins) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Common names: Snake Range bladderpod Basionym:

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Physaria valida
genusPhysaria speciesPhysaria valida (Greene) O’Kane & Al-Shehbaz Novon 12: 328. 2002. Steve L. O’Kane Jr. Common names: Strong bladderpod Basionym: Lesquerella

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Physaria fremontii
speciesPhysaria fremontii (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz Novon 12: 323. 2002. Steve L. O’Kane Jr. Common names: Fremont’s bladderpod Basionym:

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Physaria kingii subsp. kaibabensis
subspeciesPhysaria kingii subsp. kaibabensis (Rollins) O’Kane Novon 17: 380. 2007. Steve L. O’Kane Jr. Common names: Kaibab bladderpod Basionym: Lesquerella

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Physaria parviflora
genusPhysaria speciesPhysaria parviflora (Rollins) O’Kane & Al-Shehbaz Novon 12: 326. 2002. Steve L. O’Kane Jr. Common names: Picenace bladderpod Basionym: Lesquerella

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Physaria subumbellata
genusPhysaria speciesPhysaria subumbellata (Rollins) O’Kane & Al-Shehbaz Novon 12: 328. 2002. Steve L. O’Kane Jr. Common names: Parasol bladderpod Basionym: Lesquerella

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Physaria lata
genusPhysaria speciesPhysaria lata (Wooton & Standley) O’Kane & Al-Shehbaz Novon 12: 325. 2002. Steve L. O’Kane Jr. Common names: Lincoln County bladderpod Basionym:

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Physaria pulvinata
Physarieae genusPhysaria speciesPhysaria pulvinata O’Kane & Reveal Brittonia 58: 74, fig. 1. 2006. Steve L. O’Kane Jr. Common names: Cushion bladderpod Treatment

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