Difference between revisions of "Vancouveria"

C. Morren & Decaisne

Ann. Sci. Nat., Bot., sér. 2, 2: 351. 1834.

Common names: Inside-out flower
Etymology: for George Vancouver (1757-1798), English navigator and explorer
Treatment appears in FNA Volume 3.
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--><span class="statement" id="st-d0_s0" data-properties="herb duration;herb duration;herb duration;herb some measurement;herb pubescence;herb pubescence;herb pubescence;herb pubescence;herb pubescence"><b>Herbs,</b> perennial, evergreen or deciduous, 1-5 dm, glabrous, glandular-pubescent, or sparsely hairy.</span> <span class="statement" id="st-d0_s1" data-properties="rhizome size;rhizome growth form or orientation;rhizome shape;leaf quantity;shoot life cycle"><b>Rhizomes </b>extensive, creeping, nodose, producing 3 or more foliage leaves and flowering shoots per year.</span> <span class="statement" id="st-d0_s2" data-properties="stem location;stem presence"><b>Aerial </b>stems absent.</span> <span class="statement" id="st-d0_s3" data-properties="leaf position;leaf arrangement;leaf architecture"><b>Leaves </b>basal, alternate, 2-3-ternately compound (sometimes 3-5-foliolate in V. chrysantha);</span> <span class="statement" id="st-d0_s4" data-properties="petiole length or size;petiole size">petiole long, slender.</span> <span class="statement" id="st-d0_s5" data-properties="leaf-blade shape"><b>Leaf-</b>blade deltate in overall outline;</span> <span class="statement" id="st-d0_s6" data-properties="">rachis without pulvinae;</span> <span class="statement" id="st-d0_s7" data-properties="leaflet blade shape;leaflet blade shape;leaflet blade shape;leaflet blade shape;leaflet blade shape;leaflet blade shape">leaflet blades rhomboid or rounded-pentagonal to ovate to oblong, shallowly 3-lobed, margins entire to sinuate;</span> <span class="statement" id="st-d0_s8" data-properties="margin shape;margin shape;margin shape;margin architecture">venation palmate.</span> <span class="statement" id="st-d0_s9" data-properties="inflorescence position or structure subtype;raceme architecture;panicle architecture"><b>Inflorescences </b>terminal, racemes or panicles, open.</span> <span class="statement" id="st-d0_s10" data-properties="flower architecture;flower some measurement"><b>Flowers </b>3-merous, 6-14 mm;</span> <span class="statement" id="st-d0_s11" data-properties="bracteole quantity;bracteole architecture">bracteoles 6-9, sepaloid;</span> <span class="statement" id="st-d0_s12" data-properties="sepal quantity;sepal coloration;sepal coloration;sepal coloration">sepals 6, white to yellow;</span> <span class="statement" id="st-d0_s13" data-properties="petal quantity;petal coloration;petal coloration;petal coloration;petal architecture;petal prominence or shape;tip orientation">petals 6, white to yellow, hooded with tip reflexed or flat, bearing nectar;</span> <span class="statement" id="st-d0_s14" data-properties="stamen quantity">stamens 6;</span> <span class="statement" id="st-d0_s15" data-properties="anther dehiscence;flap quantity">anthers dehiscing by 2 apically hinged flaps;</span> <span class="statement" id="st-d0_s16" data-properties="pollen exine coloration or pubescence or relief">pollen exine striate;</span> <span class="statement" id="st-d0_s17" data-properties="">ovaries ellipsoid;</span> <span class="statement" id="st-d0_s18" data-properties="ovary shape;ovary placentation">placentation marginal;</span> <span class="statement" id="st-d0_s19" data-properties="style position">style lateral.</span> <span class="statement" id="st-d0_s20" data-properties="fruits follicle coloration;fruits follicle architecture or shape;fruits follicle arrangement or shape;fruits follicle dehiscence;valve quantity"><b>Fruits </b>follicles, brown, asymmetric, generally elliptic, dehiscing by 2 valves.</span> <span class="statement" id="st-d0_s21" data-properties="seed quantity;seed coloration;seed coloration;seed coloration"><b>Seeds </b>4-7, black to reddish-brown;</span> <span class="statement" id="st-d0_s22" data-properties="">aril whitish, covering ca. 1/2-2/3 of seed.</span> <span class="statement" id="st-d0_s23" data-properties="aril coloration;aril position relational;aril quantity;x chromosome quantity">x = 6.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> perennial, evergreen or deciduous, 1-5 dm, glabrous, glandular-pubescent, or sparsely hairy. <b>Rhizomes</b> extensive, creeping, nodose, producing 3 or more foliage leaves and flowering shoots per year. <b>Aerial</b> stems absent. <b>Leaves</b> basal, alternate, 2-3-ternately compound (sometimes 3-5-foliolate in V. chrysantha); petiole long, slender. <b>Leaf</b> blade deltate in overall outline; rachis without pulvinae; leaflet blades rhomboid or rounded pentagonal to ovate to oblong, shallowly 3-lobed, margins entire to sinuate; venation palmate. <b>Inflorescences</b> terminal, racemes or panicles, open. <b>Flowers</b> 3-merous, 6-14 mm; bracteoles 6-9, sepaloid; sepals 6, white to yellow; petals 6, white to yellow, hooded with tip reflexed or flat, bearing nectar; stamens 6; anthers dehiscing by 2 apically hinged flaps; pollen exine striate; ovaries ellipsoid; placentation marginal; style lateral. <b>Fruits</b> follicles, brown, asymmetric, generally elliptic, dehiscing by 2 valves. <b>Seeds</b> 4-7, black to reddish brown; aril whitish, covering ca. 1/2-2/3 of seed. <b>x</b> = 6.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=w United States
+
|distribution=w United States.
 
|discussion=<p>Species 3 (3 in the flora).</p><!--
 
|discussion=<p>Species 3 (3 in the flora).</p><!--
 
--><p>The fruits of Vancouveria are thin-walled follicles that are green or greenish brown at the time of dehiscence. The follicles dehisce by means of two valves that begin below the style and open to the base. The two valves recurve, exposing the seeds downward. In V. hexandra the follicle opens before the seeds are mature. The green seeds continue to grow and ripen in the open follicle. The appendage or aril on Vancouveria seeds has been shown to elaiosome (R. Y. Berg 1972). Ants carry the seeds to their nests and harvest the appendage as a food source.</p>
 
--><p>The fruits of Vancouveria are thin-walled follicles that are green or greenish brown at the time of dehiscence. The follicles dehisce by means of two valves that begin below the style and open to the base. The two valves recurve, exposing the seeds downward. In V. hexandra the follicle opens before the seeds are mature. The green seeds continue to grow and ripen in the open follicle. The appendage or aril on Vancouveria seeds has been shown to elaiosome (R. Y. Berg 1972). Ants carry the seeds to their nests and harvest the appendage as a food source.</p>
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|basionyms=
 
|basionyms=
 
|family=Berberidaceae
 
|family=Berberidaceae
|distribution=w United States
+
|distribution=w United States.
 
|reference=berg1972b;stearn1938a
 
|reference=berg1972b;stearn1938a
 
|publication title=Ann. Sci. Nat., Bot., sér.
 
|publication title=Ann. Sci. Nat., Bot., sér.
 
|publication year=1834
 
|publication year=1834
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_519.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_519.xml
 
|genus=Vancouveria
 
|genus=Vancouveria
|anther dehiscence=dehiscing
 
|aril coloration=whitish
 
|aril position relational=covering
 
|aril quantity=1/2;2/3
 
|bracteole architecture=sepaloid
 
|bracteole quantity=6;9
 
|flap quantity=2
 
|flower architecture=3-merous
 
|flower some measurement=6mm;14mm
 
|fruits follicle architecture or shape=asymmetric
 
|fruits follicle arrangement or shape=elliptic
 
|fruits follicle coloration=brown
 
|fruits follicle dehiscence=dehiscing
 
|herb duration=deciduous;evergreen;perennial
 
|herb pubescence=hairy;glandular-pubescent;hairy;glandular-pubescent;glabrous
 
|herb some measurement=1dm;5dm
 
|inflorescence position or structure subtype=terminal
 
|leaf architecture=compound
 
|leaf arrangement=alternate
 
|leaf position=basal
 
|leaf quantity=3
 
|leaf-blade shape=deltate
 
|leaflet blade shape=3-lobed;rounded-pentagonal;ovate
 
|margin architecture=palmate
 
|margin shape=entire;sinuate
 
|ovary placentation=marginal
 
|ovary shape=ellipsoid
 
|panicle architecture=open
 
|petal architecture=hooded
 
|petal coloration=white;yellow
 
|petal prominence or shape=flat
 
|petal quantity=6
 
|petiole length or size=long
 
|petiole size=slender
 
|pollen exine coloration or pubescence or relief=striate
 
|raceme architecture=open
 
|rhizome growth form or orientation=creeping
 
|rhizome shape=nodose
 
|rhizome size=extensive
 
|seed coloration=black;reddish-brown
 
|seed quantity=4;7
 
|sepal coloration=white;yellow
 
|sepal quantity=6
 
|shoot life cycle=flowering
 
|stamen quantity=6
 
|stem location=aerial
 
|stem presence=absent
 
|style position=lateral
 
|tip orientation=reflexed
 
|valve quantity=2
 
|x chromosome quantity=6
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Berberidaceae]]
 
-->[[Category:Treatment]][[Category:Berberidaceae]]

Revision as of 14:42, 27 July 2019

Herbs, perennial, evergreen or deciduous, 1-5 dm, glabrous, glandular-pubescent, or sparsely hairy. Rhizomes extensive, creeping, nodose, producing 3 or more foliage leaves and flowering shoots per year. Aerial stems absent. Leaves basal, alternate, 2-3-ternately compound (sometimes 3-5-foliolate in V. chrysantha); petiole long, slender. Leaf blade deltate in overall outline; rachis without pulvinae; leaflet blades rhomboid or rounded pentagonal to ovate to oblong, shallowly 3-lobed, margins entire to sinuate; venation palmate. Inflorescences terminal, racemes or panicles, open. Flowers 3-merous, 6-14 mm; bracteoles 6-9, sepaloid; sepals 6, white to yellow; petals 6, white to yellow, hooded with tip reflexed or flat, bearing nectar; stamens 6; anthers dehiscing by 2 apically hinged flaps; pollen exine striate; ovaries ellipsoid; placentation marginal; style lateral. Fruits follicles, brown, asymmetric, generally elliptic, dehiscing by 2 valves. Seeds 4-7, black to reddish brown; aril whitish, covering ca. 1/2-2/3 of seed. x = 6.

Distribution

w United States.

Discussion

Species 3 (3 in the flora).

The fruits of Vancouveria are thin-walled follicles that are green or greenish brown at the time of dehiscence. The follicles dehisce by means of two valves that begin below the style and open to the base. The two valves recurve, exposing the seeds downward. In V. hexandra the follicle opens before the seeds are mature. The green seeds continue to grow and ripen in the open follicle. The appendage or aril on Vancouveria seeds has been shown to elaiosome (R. Y. Berg 1972). Ants carry the seeds to their nests and harvest the appendage as a food source.

Key

1 Leaflet margins not conspicuously thickened; leaves falling when fruits maturing; pedicels lacking glands. Vancouveria hexandra
1 Leaflet margins conspicuously thickened; leaves persistent; pedicels stipitate-glandular. > 2
2 Petals yellow, petal apex reflexed; follicles densely stipitate-glandular. Vancouveria chrysantha
2 Petals white, sometimes lavender-tinged, petal apex not reflexed; follicles lacking glands. Vancouveria planipetala
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