Penstemon heterodoxus

A. Gray

in A. Gray et al., Syn. Fl. N. Amer. 2(1): 269. 1878. (as Pentstemon)

Common names: Sierra beardtongue
Treatment appears in FNA Volume 17. Treatment on page 207. Mentioned on page 189, 208.

Herbs. Stems decumbent, ascending, or erect, 5–65 cm, glabrous or retrorsely hairy, sometimes also glandular-pubescent distally, not glaucous. Leaves basal and cauline, not leathery, glabrous; basal and proximal cauline 5–140 × 3–20 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–5 pairs, short-petiolate or sessile, 10–60(–70) × 1–14 mm, blade oblong to oblanceolate or lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. Thyrses interrupted, cylindric, 1–12 cm, axis ± glandular-pubescent, verticillasters 1–6, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 4–35 × 1–6 mm, margins entire; peduncles and pedicels erect, ± glandular-pubescent. Flowers: calyx lobes oblong to lanceolate, 4–6 × 0.7–1.2 mm, glandular-pubescent; corolla lavender to blue, violet, or purple, essentially without nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, moderately yellow-pilose to yellow-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–5 mm diam., prominently 2-ridged abaxially; stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 0.6–0.8(–1) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 6–9 mm, included, 0.3–0.6 mm diam., tip straight to recurved, glabrous or distal 1–3 mm sparsely to moderately pilose, hairs yellow, to 0.5 mm; style 9–12 mm. Capsules 4–6 × 2.5–3.5 mm, glabrous.


Varieties 3 (3 in the flora).

Penstemon heterodoxus comprises three partially sympatric varieties centered in the Sierra Nevada of eastern California.

Selected References



1 Thyrse axes usually sparsely glandular-pubescent; n California. Penstemon heterodoxus var. shastensis
1 Thyrse axes moderately to densely glandular-pubescent; e California, wc Nevada. > 2
2 Stems 5–25 cm; basal and proximal cauline leaves 5–50 × 3–10 mm; verticillasters (1–)3; ec California, wc Nevada. Penstemon heterodoxus var. heterodoxus
2 Stems 15–35 cm; basal and proximal cauline leaves (17–)20–75 × 6–18 mm; verticillasters 2–6; se California. Penstemon heterodoxus var. cephalophorus