Difference between revisions of "Astrolepis cochisensis"

(Goodding) D. M. Benham & Windham

Amer. Fern J. 82: 57. 1992.

Basionym: Notholaena cochisensis Goodding Muhlenbergia 8: 93. 1912
Synonyms: Cheilanthes cochisensis (Goodding) Mickel Cheilanthes sinuata var. cochisensis (Goodding) Munz Notholaena sinuata var. cochisensis (Goodding) Weatherby
Treatment appears in FNA Volume 2. Treatment on page 141.
FNA>Volume Importer
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|name=Notholaena cochisensis
 
|name=Notholaena cochisensis
 
|authority=Goodding
 
|authority=Goodding
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|rank=species
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|publication_title=Muhlenbergia
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|publication_place=8: 93. 1912
 
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|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Cheilanthes cochisensis
 
|name=Cheilanthes cochisensis
 
|authority=(Goodding) Mickel
 
|authority=(Goodding) Mickel
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|rank=species
 
}} {{Treatment/ID/Synonym
 
}} {{Treatment/ID/Synonym
 
|name=Cheilanthes sinuata var. cochisensis
 
|name=Cheilanthes sinuata var. cochisensis
 
|authority=(Goodding) Munz
 
|authority=(Goodding) Munz
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|rank=variety
 
}} {{Treatment/ID/Synonym
 
}} {{Treatment/ID/Synonym
 
|name=Notholaena sinuata var. cochisensis
 
|name=Notholaena sinuata var. cochisensis
 
|authority=(Goodding) Weatherby
 
|authority=(Goodding) Weatherby
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|rank=variety
 
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}}
 
|hierarchy=Pteridaceae;Astrolepis;Astrolepis cochisensis
 
|hierarchy=Pteridaceae;Astrolepis;Astrolepis cochisensis
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|distribution=North America;Mexico.
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|distribution=Ariz.;Calif.;N.Mex.;Okla.;Tex.;Mexico.
 
|discussion=<p>Subspecies 3 (3 in the flora).</p><!--
 
|discussion=<p>Subspecies 3 (3 in the flora).</p><!--
 
--><p><i>Astrolepis cochisensis</i> is reported to be toxic to sheep (F. P. Mathews 1945). Three cytotypes that occupy different ranges and/or habitats have been identified and are treated here as subspecies. These include a sexual diploid (<i></i>subsp.<i> chihuahuensis</i>) found on calcareous substrates in the Chihuahuan Desert; an apogamous triploid (<i></i>subsp.<i> cochisensis</i>), which inhabits primarily calcareous substrates in the Sonoran, Mojavean, and western Chihuahuan deserts; and an apogamous tetraploid (<i></i>subsp.<i> arizonica</i>), which occupies primarily noncalcareous substrates in southern Arizona. Isozyme analyses suggest that <i></i>subsp.<i> cochisensis</i> is an autotriploid derivative of the diploid <i></i>subsp.<i> chihuahuensis</i> (D. M. Benham 1989). Both the isozymes and substrate preferences of <i></i>subsp.<i> arizonica</i> indicate, however, that it is not a simple autotetraploid and that other taxa remain to be discovered within the <i>Astrolepis cochisensis</i> complex.</p>
 
--><p><i>Astrolepis cochisensis</i> is reported to be toxic to sheep (F. P. Mathews 1945). Three cytotypes that occupy different ranges and/or habitats have been identified and are treated here as subspecies. These include a sexual diploid (<i></i>subsp.<i> chihuahuensis</i>) found on calcareous substrates in the Chihuahuan Desert; an apogamous triploid (<i></i>subsp.<i> cochisensis</i>), which inhabits primarily calcareous substrates in the Sonoran, Mojavean, and western Chihuahuan deserts; and an apogamous tetraploid (<i></i>subsp.<i> arizonica</i>), which occupies primarily noncalcareous substrates in southern Arizona. Isozyme analyses suggest that <i></i>subsp.<i> cochisensis</i> is an autotriploid derivative of the diploid <i></i>subsp.<i> chihuahuensis</i> (D. M. Benham 1989). Both the isozymes and substrate preferences of <i></i>subsp.<i> arizonica</i> indicate, however, that it is not a simple autotetraploid and that other taxa remain to be discovered within the <i>Astrolepis cochisensis</i> complex.</p>
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name=Astrolepis cochisensis
 
name=Astrolepis cochisensis
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|basionyms=Notholaena cochisensis
 
|basionyms=Notholaena cochisensis
 
|family=Pteridaceae
 
|family=Pteridaceae
|distribution=North America;Mexico.
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|distribution=Ariz.;Calif.;N.Mex.;Okla.;Tex.;Mexico.
 
|reference=None
 
|reference=None
 
|publication title=Amer. Fern J.
 
|publication title=Amer. Fern J.
 
|publication year=1992
 
|publication year=1992
 
|special status=
 
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/8f726806613d60c220dc4493de13607dd3150896/coarse_grained_fna_xml/V2/V2_191.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_191.xml
 
|genus=Astrolepis
 
|genus=Astrolepis
 
|species=Astrolepis cochisensis
 
|species=Astrolepis cochisensis
 
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[[Category:Revised Since Print]]

Latest revision as of 17:56, 6 November 2020

Stems compact; stem scales uniformly tan or somewhat darker near base, to 10 mm, margins ciliate-dentate to entire. Leaves 7–40 cm. Blade 1-pinnate to pinnate-pinnatifid, pinna pairs 20–50. Pinnae oblong, largest usually 4–7 mm, entire or asymmetrically lobed, lobes 1–4, broadly rounded, separated by shallow sinuses; abaxial scales completely concealing surface, ovate, usually 0.5–1 mm, ciliate with coarse marginal projections; adaxial scales sparse, deciduous, stellate to coarsely ciliate, mostly circular to elliptic, peltate, body more than 5 cells wide. Sporangia containing 32 or 64 spores.

Distribution

Ariz., Calif., N.Mex., Okla., Tex., Mexico.

Discussion

Subspecies 3 (3 in the flora).

Astrolepis cochisensis is reported to be toxic to sheep (F. P. Mathews 1945). Three cytotypes that occupy different ranges and/or habitats have been identified and are treated here as subspecies. These include a sexual diploid (subsp. chihuahuensis) found on calcareous substrates in the Chihuahuan Desert; an apogamous triploid (subsp. cochisensis), which inhabits primarily calcareous substrates in the Sonoran, Mojavean, and western Chihuahuan deserts; and an apogamous tetraploid (subsp. arizonica), which occupies primarily noncalcareous substrates in southern Arizona. Isozyme analyses suggest that subsp. cochisensis is an autotriploid derivative of the diploid subsp. chihuahuensis (D. M. Benham 1989). Both the isozymes and substrate preferences of subsp. arizonica indicate, however, that it is not a simple autotetraploid and that other taxa remain to be discovered within the Astrolepis cochisensis complex.

Selected References

None.

Key

1 Sporangia containing 64 spores; spores averaging 39– 46 µm diam.; plants of calcareous substrates in Texas, s New Mexico. Astrolepis cochisensis subsp. chihuahuensis
1 Sporangia containing 32 spores; spores averaging 59– 86 µm diam.; plants of calcareous and noncalcareous substrates in California, Arizona, Nevada, w New Mexico, extreme w Texas, Oklahoma, Utah, n Mexico. > 2
2 Spores averaging 59–70 µm diam.; plants primarily of calcareous substrates throughout Southwest. Astrolepis cochisensis subsp. cochisensis
2 Spores averaging 72–86 µm diam.; plants primarily of noncalcareous substrates in s Arizona. Astrolepis cochisensis subsp. arizonica
... more about "Astrolepis cochisensis"
Dale M. Benham +  and Michael D. Windham +
(Goodding) D. M. Benham & Windham +
Notholaena cochisensis +
Ariz. +, Calif. +, N.Mex. +, Okla. +, Tex. +  and Mexico. +
Amer. Fern J. +
Cheilanthes cochisensis +, Cheilanthes sinuata var. cochisensis +  and Notholaena sinuata var. cochisensis +
Astrolepis cochisensis +
Astrolepis +
species +