Difference between revisions of "Dicentra formosa"

(Andrews) Walpers

Repert. Bot. Syst. 1: 118. 1842.

Common names: Pacific bleeding-heart
IllustratedEndemic
Basionym: Fumaria formosa Andrews Bot. Repos. 6: plate 393. 1804
Synonyms: Dicentra saccata (Nuttall ex Torrey & A. Gray) Walpers
Treatment appears in FNA Volume 3.
FNA>Volume Importer
 
(Correct accepted name and basionym authorships, date of basionym, and deleted paragraph from the discussion about the basionym authorship.)
 
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{{Treatment/ID
 
{{Treatment/ID
 
|accepted_name=Dicentra formosa
 
|accepted_name=Dicentra formosa
|accepted_authority=(Haworth) Walpers
+
|accepted_authority=(Andrews) Walpers
 
|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=Repert. Bot. Syst.
 
|title=Repert. Bot. Syst.
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|common_names=Pacific bleeding-heart
 
|common_names=Pacific bleeding-heart
 
|special_status={{Treatment/ID/Special_status
 
|special_status={{Treatment/ID/Special_status
 +
|code=F
 +
|label=Illustrated
 +
}}{{Treatment/ID/Special_status
 
|code=E
 
|code=E
 
|label=Endemic
 
|label=Endemic
 
}}
 
}}
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Fumaria formosa
 
|name=Fumaria formosa
|authority=Haworth
+
|authority=Andrews
 +
|rank=species
 +
|publication_title=Bot. Repos.
 +
|publication_place=6: plate 393. 1804
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Dicentra saccata
 
|name=Dicentra saccata
 
|authority=(Nuttall ex Torrey & A. Gray) Walpers
 
|authority=(Nuttall ex Torrey & A. Gray) Walpers
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Fumariaceae;Dicentra;Dicentra formosa
 
|hierarchy=Fumariaceae;Dicentra;Dicentra formosa
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="plant duration;plant architecture;rhizome shape;rhizome fragility or size"><b>Plants </b>perennial, scapose, from elongate, stout rhizomes.</span> <span class="statement" id="st-d0_s1" data-properties="leaf atypical length;leaf atypical length;leaf length;leaf atypical width;leaf atypical width;leaf width"><b>Leaves </b>(15-) 25-40 (-55) × (8-) 12-20 (-35) cm;</span> <span class="statement" id="st-d0_s2" data-properties="order quantity">blade with 3-5 orders of leaflets and lobes;</span> <span class="statement" id="st-d0_s3" data-properties="abaxial surface position;abaxial surface pubescence">abaxial surface and sometimes adaxial surface glaucous;</span> <span class="statement" id="st-d0_s4" data-properties="penultimate lobe shape;penultimate lobe position or shape;penultimate lobe shape;apex atypical length;apex atypical length;apex length;apex atypical width;apex atypical width;apex width">penultimate lobes oblong, distal ones usually coarsely 3-toothed at apex, (4-) 10-20 (-50) × (1.5-) 3-4 (-8) mm.</span> <span class="statement" id="st-d0_s5" data-properties="inflorescence arrangement;inflorescence architecture"><b>Inflorescences </b>paniculate, 2-30-flowered, usually exceeding leaves;</span> <span class="statement" id="st-d0_s6" data-properties="beak shape;beak atypical length;beak length;beak width;apex shape">bracts linear-lanceolate, 4-7 (-12) × 1-2 mm, apex acuminate.</span> <span class="statement" id="st-d0_s7" data-properties="flower orientation"><b>Flowers </b>pendent;</span> <span class="statement" id="st-d0_s8" data-properties="sepal shape;sepal shape;sepal shape;sepal shape;sepal length;sepal width">sepals lanceolate to ovate or nearly round, 2-7 × 2-3 mm;</span> <span class="statement" id="st-d0_s9" data-properties="petal coloration;petal coloration;petal coloration;petal coloration;petal coloration;petal coloration;petal coloration">petals rose-purple, pink, cream, or pale-yellow, rarely white;</span> <span class="statement" id="st-d0_s10" data-properties="outer petal atypical length;outer petal atypical length;outer petal length;outer petal width;portion orientation;portion some measurement">outer petals (12-) 16-19 (-24) × 3-6 mm, reflexed portion 2-5 mm;</span> <span class="statement" id="st-d0_s11" data-properties="inner petal atypical some measurement;inner petal atypical some measurement;inner petal some measurement;blade width;claw shape;claw shape;claw shape;claw atypical length;claw length;claw width;crest diameter">inner petals (12-) 15-18 (-22) mm, blade 2-4 mm wide, claw linear-elliptic to linear-lanceolate, 7-10 (-12) × 1-2 mm, crest 1-2 mm diam., exceeding apex by 1-2 mm;</span> <span class="statement" id="st-d0_s12" data-properties="bundle filament fusion;portion some measurement">filaments of each bundle connate from base to shortly below anthers except for a 2-3 mm portion of median filament just above base;</span> <span class="statement" id="st-d0_s13" data-properties="portion fusion;median filament fusion">nectariferous tissue borne along distinct portion of median filament;</span> <span class="statement" id="st-d0_s14" data-properties="style some measurement">style 3-9 mm;</span> <span class="statement" id="st-d0_s15" data-properties="stigma shape;stigma shape">stigma rhomboid, 2-horned.</span> <span class="statement" id="st-d0_s16" data-properties="capsule shape;capsule diameter"><b>Capsules </b>oblong, 4-5 mm diam.</span> <span class="statement" id="st-d0_s17" data-properties="seed shape;seed diameter;seed architecture or coloration or relief;elaiosome presence"><b>Seeds </b>reniform, ca. 2 mm diam., finely reticulate, elaiosome present.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>perennial, scapose, from elongate, stout rhizomes. <b>Leaves</b> (15-)25-40(-55) × (8-)12-20(-35) cm; blade with 3-5 orders of leaflets and lobes; abaxial surface and sometimes adaxial surface glaucous; penultimate lobes oblong, distal ones usually coarsely 3-toothed at apex, (4-)10-20(-50) × (1.5-)3-4(-8) mm. <b>Inflorescences</b> paniculate, 2-30-flowered, usually exceeding leaves; bracts linear-lanceolate, 4-7(-12) × 1-2 mm, apex acuminate. <b>Flowers</b> pendent; sepals lanceolate to ovate or nearly round, 2-7 × 2-3 mm; petals rose-purple, pink, cream, or pale yellow, rarely white; outer petals (12-)16-19(-24) × 3-6 mm, reflexed portion 2-5 mm; inner petals (12-)15-18(-22) mm, blade 2-4 mm wide, claw linear-elliptic to linear-lanceolate, 7-10(-12) × 1-2 mm, crest 1-2 mm diam., exceeding apex by 1-2 mm; filaments of each bundle connate from base to shortly below anthers except for a 2-3 mm portion of median filament just above base; nectariferous tissue borne along distinct portion of median filament; style 3-9 mm; stigma rhomboid, 2-horned. <b>Capsules</b> oblong, 4-5 mm diam. <b>Seeds</b> reniform, ca. 2 mm diam., finely reticulate, elaiosome present.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=w North America
+
|distribution=B.C.;Calif.;Oreg.;Wash.
 
|discussion=<p>Subspecies 2 (2 in the flora).</p><!--
 
|discussion=<p>Subspecies 2 (2 in the flora).</p><!--
--><p>Andrews has been cited almost universally as the author of Fumaria formosa. However, Haworth's authorship of the sixth volume of Andrews' Botanists' Repository (in which this species was originally described) generally has been overlooked, and it was actually Haworth who first delineated F. formosa (W. T. Stearn 1944).</p><!--
+
--><p>Early attempts to cross <i>Dicentra formosa</i> with <i>D. eximia</i> (2n = 16) failed, possibly because the <i>D. formosa</i> parents were tetraploids. Several later hybrids between the two species received plant patents and have become widely marketed throughout the flora area and elsewhere (K. R. Stern 1961, 1968; K. R. Stern and M. Ownbey 1971).</p><!--
--><p>Early attempts to cross Dicentra formosa with D. eximia (2n = 16) failed, possibly because the D. formosa parents were tetraploids. Several later hybrids between the two species received plant patents and have become widely marketed throughout the flora area and elsewhere (K. R. Stern 1961, 1968; K. R. Stern and M. Ownbey 1971).</p><!--
+
--><p>Both subspecies, as well as hybrids between them and <i>Dicentra eximia</i>, are widely cultivated.</p><!--
--><p>Both subspecies, as well as hybrids between them and Dicentra eximia, are widely cultivated.</p><!--
+
--><p>The Skagit used a decoction of the roots of <i>Dicentra formosa</i> to expel worms; they chewed raw roots for toothaches (D. E. Moerman 1986, species not indicated).</p>
--><p>The Skagit used a decoction of the roots of Dicentra formosa to expel worms; they chewed raw roots for toothaches (D. E. Moerman 1986, species not indicated).</p>
 
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Dicentra formosa
 
name=Dicentra formosa
|author=
+
|authority=(Andrews) Walpers
|authority=(Haworth) Walpers
 
 
|rank=species
 
|rank=species
 
|parent rank=genus
 
|parent rank=genus
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|basionyms=Fumaria formosa
 
|basionyms=Fumaria formosa
 
|family=Fumariaceae
 
|family=Fumariaceae
|distribution=w North America
+
|distribution=B.C.;Calif.;Oreg.;Wash.
 
|reference=None
 
|reference=None
 
|publication title=Repert. Bot. Syst.
 
|publication title=Repert. Bot. Syst.
 
|publication year=1842
 
|publication year=1842
|special status=Endemic
+
|special status=Illustrated;Endemic
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_1080.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V3/V3_1080.xml
 
|genus=Dicentra
 
|genus=Dicentra
 
|species=Dicentra formosa
 
|species=Dicentra formosa
|abaxial surface position=adaxial
 
|abaxial surface pubescence=glaucous
 
|apex atypical length=20mm;50mm
 
|apex atypical width=4mm;8mm
 
|apex length=10mm;20mm
 
|apex shape=acuminate
 
|apex width=3mm;4mm
 
|beak atypical length=7mm;12mm
 
|beak length=4mm;7mm
 
|beak shape=linear-lanceolate
 
|beak width=1mm;2mm
 
|blade width=2mm;4mm
 
|bundle filament fusion=connate
 
|capsule diameter=4mm;5mm
 
|capsule shape=oblong
 
|claw atypical length=10mm;12mm
 
|claw length=7mm;10mm
 
|claw shape=linear-elliptic;linear-lanceolate
 
|claw width=1mm;2mm
 
|crest diameter=1mm;2mm
 
|elaiosome presence=absent
 
|flower orientation=pendent
 
|inflorescence architecture=2-30-flowered
 
|inflorescence arrangement=paniculate
 
|inner petal atypical some measurement=18mm;22mm
 
|inner petal some measurement=15mm;18mm
 
|leaf atypical length=40cm;55cm
 
|leaf atypical width=20cm;35cm
 
|leaf length=25cm;40cm
 
|leaf width=12cm;20cm
 
|median filament fusion=distinct
 
|order quantity=3;5
 
|outer petal atypical length=19mm;24mm
 
|outer petal length=16mm;19mm
 
|outer petal width=3mm;6mm
 
|penultimate lobe position or shape=distal
 
|penultimate lobe shape=3-toothed;oblong
 
|petal coloration=white;pale-yellow;cream;pale-yellow;cream;pink;rose-purple
 
|plant architecture=scapose
 
|plant duration=perennial
 
|portion fusion=distinct
 
|portion orientation=reflexed
 
|portion some measurement=2mm;3mm
 
|rhizome fragility or size=stout
 
|rhizome shape=elongate
 
|seed architecture or coloration or relief=reticulate
 
|seed diameter=2
 
|seed shape=reniform
 
|sepal length=2mm;7mm
 
|sepal shape=lanceolate;ovate or nearly round
 
|sepal width=2mm;3mm
 
|stigma shape=2-horned;rhomboid
 
|style some measurement=3mm;9mm
 
 
}}<!--
 
}}<!--
  
-->[[Category:Treatment]][[Category:Dicentra]]
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-->
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 +
[[Category:Treatment]]
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[[Category:Dicentra]]
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[[Category:Revised Since Print]]

Latest revision as of 12:01, 8 May 2023

Plants perennial, scapose, from elongate, stout rhizomes. Leaves (15-)25-40(-55) × (8-)12-20(-35) cm; blade with 3-5 orders of leaflets and lobes; abaxial surface and sometimes adaxial surface glaucous; penultimate lobes oblong, distal ones usually coarsely 3-toothed at apex, (4-)10-20(-50) × (1.5-)3-4(-8) mm. Inflorescences paniculate, 2-30-flowered, usually exceeding leaves; bracts linear-lanceolate, 4-7(-12) × 1-2 mm, apex acuminate. Flowers pendent; sepals lanceolate to ovate or nearly round, 2-7 × 2-3 mm; petals rose-purple, pink, cream, or pale yellow, rarely white; outer petals (12-)16-19(-24) × 3-6 mm, reflexed portion 2-5 mm; inner petals (12-)15-18(-22) mm, blade 2-4 mm wide, claw linear-elliptic to linear-lanceolate, 7-10(-12) × 1-2 mm, crest 1-2 mm diam., exceeding apex by 1-2 mm; filaments of each bundle connate from base to shortly below anthers except for a 2-3 mm portion of median filament just above base; nectariferous tissue borne along distinct portion of median filament; style 3-9 mm; stigma rhomboid, 2-horned. Capsules oblong, 4-5 mm diam. Seeds reniform, ca. 2 mm diam., finely reticulate, elaiosome present.

Discussion

Subspecies 2 (2 in the flora).

Early attempts to cross Dicentra formosa with D. eximia (2n = 16) failed, possibly because the D. formosa parents were tetraploids. Several later hybrids between the two species received plant patents and have become widely marketed throughout the flora area and elsewhere (K. R. Stern 1961, 1968; K. R. Stern and M. Ownbey 1971).

Both subspecies, as well as hybrids between them and Dicentra eximia, are widely cultivated.

The Skagit used a decoction of the roots of Dicentra formosa to expel worms; they chewed raw roots for toothaches (D. E. Moerman 1986, species not indicated).

Selected References

None.

Key

1 Petals rose-purple to pink, rarely white; leaf blades adaxially not glaucous (rarely glaucescent). Dicentra formosa subsp. formosa
1 Petals cream-colored or rarely pale yellow, rose-tipped; leaf blades adaxially distinctly glaucous. Dicentra formosa subsp. oregana