Physaria douglasii

(S. Watson) O’Kane & Al-Shehbaz

Novon 12: 322. 2002.

Common names: Douglas’s bladderpod
Basionym: Lesquerella douglasii S. Watson Proc. Amer. Acad. Arts 23: 255. 1888
Treatment appears in FNA Volume 7. Treatment on page 634. Mentioned on page 623.

Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Stems simple from base, erect, (usually unbranched), to 4.5 dm. Basal leaves: blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. Cauline leaves similar to basal, blade narrowly linear or, sometimes, orbicular. Racemes loose (lax). Fruiting pedicels (recurved, straight, curved, or sigmoid), 6–20 mm. Flowers: sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. Fruits obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. Seeds flattened. 2n = 10, 30.


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B.C., Idaho, Mont., Oreg., Wash.


Subspecies 2 (2 in the flora).

Selected References



1 Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington. Physaria douglasii subsp. douglasii
1 Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington. Physaria douglasii subsp. tuplashensis