Transberingia bursifolia

(de Candolle) Al-Shehbaz & O’Kane

Novon 13: 396. 2003.

Basionym: Nasturtium bursifolium de Candolle Syst. Nat. 2: 194. 1821
Synonyms: Arabidopsis bursifolia (de Candolle) Botschantzev Beringia bursifolia (de Candolle) R. A. Price, Al-Shehbaz & O’Kane
Treatment appears in FNA Volume 7. Treatment on page 456.
Revision as of 23:18, 16 December 2019 by FNA>Volume Importer

Plants densely hirsute basally, trichomes primarily simple, to 1 mm, these often mixed with smaller, forked, and/or dendritic ones, glabrescent or sparsely pubescent distally. Stems simple or few to several from caudex, (0.5–)1–4.5(–6.5) dm. Basal leaves: petiole 0.3–3 cm, flattened at base, ciliate; blade oblanceolate to obovate, or linear-lanceolate, (0.5–)1.2–6.5(–10) cm × (2–)3–8(–17) mm, base cuneate to attenuate, apex obtuse to subacute, surfaces pubescent, trichomes usually branched. Cauline leaves: blade lanceolate to linear-lanceolate or oblong, (0.5–)1–3(–4.2) cm × (1–)2.5–7(–10) mm, base strongly sagittate. Fruiting pedicels not appressed to rachis, straight, (3–)5–14(–18) mm, glabrous. Flowers: sepals 1.2–2.5 × 0.6–1 mm, sparsely to densely pubescent; petals (2.5–)3–4(–4.5) × (0.8–)1–1.5 mm; filaments 2–2.5 mm; anthers 0.4–0.5 mm. Fruits (1.5–)2–3.3(–4) cm × (0.7–)1–1.5(–2) mm; style slender, 0.2–0.8(–1) mm. Seeds 0.8–1 × 0.4–0.5 mm.

Distribution

V7 696-distribution-map.gif

n, w North America, e Asia (Russian Far East).

Discussion

Subspecies 2 (2 in the flora).

R. C. Rollins (1993) treated subspp. bursifolia and virgata as distinct species of Halimolobos, but the alleged differences (stems branched basally versus distally, plants cespitose versus non-cespitose, fruits slightly compressed versus terete) do not hold, and the only reliable difference between the two taxa is in the type of trichomes. Therefore, a subspecific, rather than specific, rank is adopted here.

Selected References

None.

Key

1 Plants glabrous or sparsely pubescent on stems distally and on rachises, trichomes simple, spreading. Transberingia bursifolia subsp. bursifolia
1 Plants usually pubescent, rarely glabrescent, on stems distally and on rachises, trichomes branched, subappressed, these sometimes mixed with simple ones. Transberingia bursifolia subsp. virgata