Codriophorus sect. Codriophorus

Synonyms: Racomitrium sect. Papillosa Chevallier Trichostomum sect. Stenotrichum
Treatment appears in FNA Volume 27. Treatment on page 296.

Stem sparingly or freely irregularly, dichotomously or fasciculately branched, sometimes filiform and almost unbranched, absent short, lateral tuft-like branchlets. Leaves lingulate, elliptic, broadly ovate, ovate- or oblong-lanceolate to broadly or occasionally narrowly lanceolate; margins 1–2(–4)-stratose, recurved, reflexed, incurved or plane on both sides; apices narrowly to broadly rounded-obtuse or long acuminate to acute, muticous or very rarely shortly hyaline-tipped, entire or sharply to bluntly, remotely irregularly toothed, erose-dentate, sinuate to cristate in the upper 1/5–1/4; costa strong and broad, ceasing in mid leaf to subpercurrent, in transverse-section 2–4-stratose in the distal and medial parts, weakly to strongly convex, reniform, crescent-shaped to semi-terete abaxially, flat or convex on the adaxial side, in the basal part 3–6(–7)-layered, situated at the bottom of a shallow or deeper, usually wide-angled groove, lunate, reniform or rectangular and distinctly flattened, strongly convex on the abaxial side; basal marginal laminal cells not differentiated; distal laminal cells 1- to variously 2-stratose, irregularly rounded-quadrate, subquadrate, oval to rounded-hexagonal, strongly papillose to smooth, straight-walled in the distal part.


North America, South America (Colombia), Eurasia, n Africa, Atlantic Islands (Azores, Canary Islands, Iceland, Madeira).


Species 9 (6 in the flora).

Members of sect. Codriophorus are recognized by their lingulate, broadly ovate to ovate- or oblong-lanceolate leaves, isodiametric and straight-walled distal laminal cells, and costa that is broad and robust, usually over 100 µm wide at the base. The costa is situated in a shallow, wide-angled groove and is mostly 2–4-stratose in the distal and medial parts, 3–6(–7)-stratose in the basal part, and anatomically strongly differentiated.

Selected References



1 Laminal cells smooth or very slightly papillose on young leaves; leaf margins entire or with a few irregular teeth at the apex. Codriophorus depressus
1 Laminal cells distinctly papillose; leaf margins remotely irregularly toothed, erose-dentate, sinuate to cristate in the distal 1/5-1/4, if entire then broadly rounded-obtuse > 2
2 Leaves lanceolate; apex irregularly bluntly erose-dentate; costa markedly convex and crescent-shaped abaxially. Codriophorus ryszardii
2 Leaves lingulate, elliptical to broadly ovate or ovate-lanceolate; apex broadly rounded-obtuse to acute, irregularly dentate to entire > 3
3 Costa subpercurrent, entire and only shortly forked at the tip > 4
3 Costa extending 1/2-3/4 way up the leaf, distinctly spurred > 5
4 Laminal cells strongly papillose, 1- or very rarely 2-stratose; leaves unbordered. Codriophorus acicularis
4 Laminal cells indistinctly papillose over the cell walls, always variously 2-stratose in the distal part; leaves distinctly bordered with fleshy and broad limbidia. Codriophorus norrisii
5 Leaf apex broadly rounded, flat to subcucullate, always entire or sometimes sinuate to remotely bluntly erose-dentate. Codriophorus mollis
5 Leaf apex broadly rounded-obtuse to subacute, often tubular, strongly dentate or very rarely subentire. Codriophorus aduncoides
Halina Bednarek-Ochyra +  and Ryszard Ochyra +
P. Beauvois +
North America +, South America (Colombia) +, Eurasia +, n Africa +, Atlantic Islands (Azores +, Canary Islands +, Iceland +  and Madeira). +
Mém. Soc. Linn. Paris +
Racomitrium sect. Papillosa +  and Trichostomum sect. Stenotrichum +
Codriophorus sect. Codriophorus +
Codriophorus +
section +