Mem. Calif. Acad. Sci. 1: 16. 1868,.
Plants robust, tufted, glossy, green to dark green. Stems erect or repent, to 5 cm. Leaves crispate-contorted when dry, narrowly acuminate, 4–6 mm; margins coarsely serrate distally, recurved on one or both sides proximally; apex plane or with erect margins but not cucullate. Specialized asexual reproduction absent. Seta 1–2(–3) per perichaetium, 4–10 mm. Capsule cylindric, 2.5 mm, smooth to weakly striate-ribbed when dry; peristome teeth divided into filiform segments, densely papillose. Calyptra lobes 1/2 or more length of calyptra.
Phenology: Capsules mature Mar–Sep.
Habitat: Limestone, basalt, and other rocks, and concrete, rarely soil, logs, and charred wood, open sites, especially along rivers
Elevation: low to moderate elevations (0-1400 m)
B.C., Calif., Idaho, Mont., Oreg., Wash., Asia.
Ptychomitrium gardneri are robust glossy plants easy to recognize by their green to dark green color, serrate, acuminate leaves, and narrow lobes of the deeply divided calyptra. The lobes of dry mature calyptrae often spread outward like the spokes of a wheel. The leaves are much longer and more narrowly acuminate than those of the somewhat similar P. serratum; the ranges of the latter and of P. gardneri do not overlap.