Difference between revisions of "Coryphantha macromeris"

(Engelmann) Lemaire

Cactées, 35. 1868.

Common names: Nipple beehive cactus
Basionym: Mammillaria macromeris Engelmann
Synonyms: Coryphantha macromeris var. runyonii (Britton & Rose) L. D. BensonCoryphantha macromeris WerdermannCoryphantha pirtlei Britton & RoseCoryphantha runyonii (Engelmann) BackebergLepidocoryphantha macromeris unknown
Treatment appears in FNA Volume 4. Treatment on page 224. Mentioned on page 219, 220, 222, 225, 236.
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--><span class="statement" id="st-d0_s0" data-properties="plant architecture;plant diameter;mat position;mound position;mound shape;branch life cycle;branch prominence;branch relief;spine orientation"><b>Plants </b>profusely branched, ultimately forming low mats or hemispheric mounds to 100 cm diam., immature branches sometimes predominant, conspicuously tuberculate with projecting spines.</span> <span class="statement" id="st-d0_s1" data-properties="root texture;root size;root architecture"><b>Roots </b>± succulent in largest, often massive and difficult to excavate.</span> <span class="statement" id="st-d0_s2" data-properties="stem location;portion location;portion prominence;portion shape;portion shape;portion shape;portion shape;portion shape;portion length;portion atypical width;portion atypical width;portion width;branch life cycle"><b>Stems </b>deep-seated, aerial portion conspicuous, hemispheric to short cylindric, shape sometimes obscured by profusion of immature branches, 5–23 × (1.5–) 4–8 (–13.5) cm;</span> <span class="statement" id="st-d0_s3" data-properties="tubercle size;tubercle prominence;tubercle atypical length;tubercle atypical length;tubercle length;tubercle width;tubercle texture;tubercle texture">tubercles unusually large, conspicuous, (10–) 15–38 (–45) × 6–15 mm, ± flaccid or flabby;</span> <span class="statement" id="st-d0_s4" data-properties="areolar gland prominence">areolar glands seasonally conspicuous;</span> <span class="statement" id="st-d0_s5" data-properties="areolar groove height or length or size;distance count">areolar grooves short, extending 1/2–3/4 distance from spines toward tubercle axils;</span> <span class="statement" id="st-d0_s6" data-properties="parenchyma coating">parenchyma mucilaginous;</span> <span class="statement" id="st-d0_s7" data-properties="pith size or width;pith count">pith narrow, ca. 1/10 of lesser stem diam.;</span> <span class="statement" id="st-d0_s8" data-properties="medullary system architecture;medullary system count">medullary vascular system absent.</span> <span class="statement" id="st-d0_s9" data-properties="spine atypical count;spine count"><b>Spines </b>7–21 [–55] per areole;</span> <span class="statement" id="st-d0_s10" data-properties="spine arrangement;spine atypical count;spine atypical count;spine count;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine atypical some measurement;spine atypical some measurement;spine some measurement">radial spines (3–) 9–15 (–18) per areole, white, gray, tan, or brown, (9–) 16–25 (–50) mm;</span> <span class="statement" id="st-d0_s11" data-properties="subcentral spine count">subcentral spines 2–3 in adaxial part of areole;</span> <span class="statement" id="st-d0_s12" data-properties="central spine atypical count;central spine count;central spine coloration;central spine coloration;central spine coloration;abaxial central spine orientation;abaxial central spine orientation;other orientation;other orientation;other course;other arrangement or shape;other prominence or shape;other architecture;other shape;side count;abaxial spine fragility;abaxial spine atypical some measurement;abaxial spine atypical some measurement;abaxial spine some measurement;abaxial spine variability;abaxial spine length">central spines (1–) 3–8 per areole, (tan to) pale gray to black, abaxial central spine porrect or descending, others weakly appressed or ± projecting spines, slightly curved, usually angular in cross-section (terete), sometimes flat and grooved on 1 side and rounded on the other, often slightly flexible, (15–) 25–35 (–55) mm, all equal or abaxial spine longest.</span> <span class="statement" id="st-d0_s13" data-properties="flower position;flower position;flower atypical length;flower length;flower atypical width;flower width"><b>Flowers </b>apical or nearly so, 30–50 (–60) × (30–) 40–70 mm;</span> <span class="statement" id="st-d0_s14" data-properties="outer tepal shape">outer tepals heavily fringed;</span> <span class="statement" id="st-d0_s15" data-properties="inner tepal count;inner tepal coloration;inner tepal coloration;inner tepal length;inner tepal width;midstripe coloration;margin coloration">inner tepals 20–25 per flower, bright-rose-pink or magenta, often with darker midstripes and paler margins, 30–40 × 4.5–6 mm;</span> <span class="statement" id="st-d0_s16" data-properties="outer filament coloration">outer filaments greenish white throughout or distally purplish-pink;</span> <span class="statement" id="st-d0_s17" data-properties="anther coloration">anthers bright-yellow;</span> <span class="statement" id="st-d0_s18" data-properties="stigma lobe count;stigma lobe coloration;stigma lobe coloration;stigma lobe some measurement">stigma lobes 6–13, white or pale-yellow, 3–6 mm.</span> <span class="statement" id="st-d0_s19" data-properties="fruit coloration;fruit shape;fruit shape;fruit shape;fruit shape;fruit atypical length;fruit atypical length;fruit length;fruit width"><b>Fruits </b>dark green, ovoid to obpyriform or ellipsoid, (10–) 13–25 (–30) × 12–18 mm;</span> <span class="statement" id="st-d0_s20" data-properties="floral remnant duration">floral remnant strongly persistent.</span> <span class="statement" id="st-d0_s21" data-properties=""><b>Seeds </b>reddish-brown, ± comma-shaped to spheric, 1.2–1.5 mm, finely and weakly raised-reticulate.</span> <span class="statement" id="st-d0_s22" data-properties="seed coloration;seed shape;seed shape;seed shape;seed some measurement;seed architecture or coloration or relief;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>profusely branched, ultimately forming low mats or hemispheric mounds to 100 cm diam., immature branches sometimes predominant, conspicuously tuberculate with projecting spines. <b>Roots</b> ± succulent in largest, often massive and difficult to excavate. <b>Stems</b> deep-seated, aerial portion conspicuous, hemispheric to short cylindric, shape sometimes obscured by profusion of immature branches, 5–23 × (1.5–)4–8(–13.5) cm; tubercles unusually large, conspicuous, (10–)15–38(–45) × 6–15 mm, ± flaccid or flabby; areolar glands seasonally conspicuous; areolar grooves short, extending 1/2–3/4 distance from spines toward tubercle axils; parenchyma mucilaginous; pith narrow, ca. 1/10 of lesser stem diam.; medullary vascular system absent. <b>Spines</b> 7–21[–55] per areole; radial spines (3–)9–15(–18) per areole, white, gray, tan, or brown, (9–)16–25(–50) mm; subcentral spines 2–3 in adaxial part of areole; central spines (1–)3–8 per areole, (tan to) pale gray to black, abaxial central spine porrect or descending, others weakly appressed or ± projecting spines, slightly curved, usually angular in cross section (terete), sometimes flat and grooved on 1 side and rounded on the other, often slightly flexible, (15–)25–35(–55) mm, all equal or abaxial spine longest. <b>Flowers</b> apical or nearly so, 30–50(–60) × (30–)40–70 mm; outer tepals heavily fringed; inner tepals 20–25 per flower, bright rose-pink or magenta, often with darker midstripes and paler margins, 30–40 × 4.5–6 mm; outer filaments greenish white throughout or distally purplish pink; anthers bright yellow; stigma lobes 6–13, white or pale yellow, 3–6 mm. <b>Fruits</b> dark green, ovoid to obpyriform or ellipsoid, (10–)13–25(–30) × 12–18 mm; floral remnant strongly persistent. <b>Seeds</b> reddish brown, ± comma-shaped to spheric, 1.2–1.5 mm, finely and weakly raised-reticulate. <b>2n</b> = 22.</span><!--
  
 
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|phenology=Flowering Feb–Sep; fruiting (May-)Aug–Dec.
 
|habitat=Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes
 
|habitat=Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes
 
|elevation=30-1700(-2000) m
 
|elevation=30-1700(-2000) m
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|basionyms=Mammillaria macromeris
 
|basionyms=Mammillaria macromeris
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering Feb–Sep; fruiting (May-)Aug–Dec.
 
|habitat=Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes
 
|habitat=Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes
 
|elevation=30-1700(-2000) m
 
|elevation=30-1700(-2000) m
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|publication year=1868
 
|publication year=1868
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_412.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_412.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Coryphantha
 
|genus=Coryphantha
 
|species=Coryphantha macromeris
 
|species=Coryphantha macromeris
|2n chromosome count=22
 
|abaxial central spine orientation=descending;porrect
 
|abaxial spine atypical some measurement=35mm;55mm
 
|abaxial spine fragility=flexible
 
|abaxial spine length=longest
 
|abaxial spine some measurement=25mm;35mm
 
|abaxial spine variability=equal
 
|anther coloration=bright-yellow
 
|areolar gland prominence=conspicuous
 
|areolar groove height or length or size=short
 
|branch life cycle=immature;immature
 
|branch prominence=predominant
 
|branch relief=tuberculate
 
|central spine atypical count=1;3
 
|central spine coloration=pale gray;black
 
|central spine count=3;8
 
|distance count=1/2;3/4
 
|floral remnant duration=persistent
 
|flower atypical length=50mm;60mm
 
|flower atypical width=30mm;40mm
 
|flower length=30mm;50mm
 
|flower position=nearly;apical
 
|flower width=40mm;70mm
 
|fruit atypical length=25mm;30mm
 
|fruit coloration=dark green
 
|fruit length=13mm;25mm
 
|fruit shape=ovoid;obpyriform or ellipsoid
 
|fruit width=12mm;18mm
 
|inner tepal coloration=magenta;bright-rose-pink
 
|inner tepal count=20;25
 
|inner tepal length=30mm;40mm
 
|inner tepal width=4.5mm;6mm
 
|margin coloration=paler
 
|mat position=low
 
|medullary system architecture=vascular
 
|medullary system count=absent
 
|midstripe coloration=darker
 
|mound position=low
 
|mound shape=hemispheric
 
|other architecture=grooved
 
|other arrangement or shape=angular
 
|other course=curved
 
|other orientation=projecting;appressed
 
|other prominence or shape=flat
 
|other shape=rounded
 
|outer filament coloration=greenish white throughout or distally purplish-pink
 
|outer tepal shape=fringed
 
|parenchyma coating=mucilaginous
 
|pith count=1/10
 
|pith size or width=narrow
 
|plant architecture=branched
 
|plant diameter=0cm;100cm
 
|portion atypical width=8cm;13.5cm
 
|portion length=5cm;23cm
 
|portion location=aerial
 
|portion prominence=conspicuous
 
|portion shape=obscured;hemispheric;short cylindric
 
|portion width=4cm;8cm
 
|root architecture=excavate
 
|root size=massive
 
|root texture=succulent
 
|seed architecture or coloration or relief=raised-reticulate
 
|seed coloration=reddish-brown
 
|seed shape=less comma-shaped;spheric
 
|seed some measurement=1.2mm;1.5mm
 
|side count=1
 
|spine arrangement=radial
 
|spine atypical count=15;18
 
|spine atypical some measurement=25mm;50mm
 
|spine coloration=brown;tan;brown;tan;gray;white
 
|spine count=9;15
 
|spine orientation=projecting
 
|spine some measurement=16mm;25mm
 
|stem location=deep-seated
 
|stigma lobe coloration=pale-yellow;white
 
|stigma lobe count=6;13
 
|stigma lobe some measurement=3mm;6mm
 
|subcentral spine count=2;3
 
|tubercle atypical length=38mm;45mm
 
|tubercle length=15mm;38mm
 
|tubercle prominence=conspicuous
 
|tubercle size=large
 
|tubercle texture=flabby;flaccid
 
|tubercle width=6mm;15mm
 
 
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-->[[Category:Treatment]][[Category:Coryphantha]]
 
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Revision as of 15:10, 27 July 2019

Plants profusely branched, ultimately forming low mats or hemispheric mounds to 100 cm diam., immature branches sometimes predominant, conspicuously tuberculate with projecting spines. Roots ± succulent in largest, often massive and difficult to excavate. Stems deep-seated, aerial portion conspicuous, hemispheric to short cylindric, shape sometimes obscured by profusion of immature branches, 5–23 × (1.5–)4–8(–13.5) cm; tubercles unusually large, conspicuous, (10–)15–38(–45) × 6–15 mm, ± flaccid or flabby; areolar glands seasonally conspicuous; areolar grooves short, extending 1/2–3/4 distance from spines toward tubercle axils; parenchyma mucilaginous; pith narrow, ca. 1/10 of lesser stem diam.; medullary vascular system absent. Spines 7–21[–55] per areole; radial spines (3–)9–15(–18) per areole, white, gray, tan, or brown, (9–)16–25(–50) mm; subcentral spines 2–3 in adaxial part of areole; central spines (1–)3–8 per areole, (tan to) pale gray to black, abaxial central spine porrect or descending, others weakly appressed or ± projecting spines, slightly curved, usually angular in cross section (terete), sometimes flat and grooved on 1 side and rounded on the other, often slightly flexible, (15–)25–35(–55) mm, all equal or abaxial spine longest. Flowers apical or nearly so, 30–50(–60) × (30–)40–70 mm; outer tepals heavily fringed; inner tepals 20–25 per flower, bright rose-pink or magenta, often with darker midstripes and paler margins, 30–40 × 4.5–6 mm; outer filaments greenish white throughout or distally purplish pink; anthers bright yellow; stigma lobes 6–13, white or pale yellow, 3–6 mm. Fruits dark green, ovoid to obpyriform or ellipsoid, (10–)13–25(–30) × 12–18 mm; floral remnant strongly persistent. Seeds reddish brown, ± comma-shaped to spheric, 1.2–1.5 mm, finely and weakly raised-reticulate. 2n = 22.


Phenology: Flowering Feb–Sep; fruiting (May-)Aug–Dec.
Habitat: Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes
Elevation: 30-1700(-2000) m

Distribution

V4 412-distribution-map.gif

N.Mex., Tex., Mexico (Chihuahua, Coahuila, Durango, Zacatecas).

Discussion

Southern Texas populations of Coryphantha macromeris contain atypical individuals with proliferating small stems and shorter (stunted?) spines. Sexually mature stems branch from the tubercle axils, and the whole shoot becomes covered by immature branchlets. The immature branchlets proliferate profusely and asymmetrically faster than they can reach sexual maturity, obscuring the underlying symmetry of mature stems and forming irregular, asymmetric mounds. Such plants are the basis for C. macromeris var. runyonii (Britton & Rose) L. D. Benson, but they do not grow in pure populations. Therefore the name runyonii can not be used at varietal rank without including plants morphologically similar to typical C. macromeris from the Chihuahuan Desert.

Stunted or immature Coryphantha macromeris are variable, keying to Coryphantha with some difficulty, often having only 5–7 radial spines and lacking central spines. The strongly mucilaginous cortex is a useful field mark; even small slices of living tubercle tissue are visibly and tangibly slimy.

Selected References

None.

Lower Taxa

None.
... more about "Coryphantha macromeris"
Allan D. Zimmerman +  and Bruce D. Parfitt +
(Engelmann) Lemaire +
Mammillaria macromeris +
Nipple beehive cactus +
N.Mex. +, Tex. +, Mexico (Chihuahua +, Coahuila +, Durango +  and Zacatecas). +
30-1700(-2000) m +
Chihuahuan desert scrub, Tamaulipan thorn scrub, nearly all substrates including nearly pure gypsum, gravelly soils, usually sandy alluvium or clay, rarely crevices or steep slopes +
Flowering Feb–Sep +  and fruiting (May-)Aug–Dec. +
Coryphantha macromeris var. runyonii +, Coryphantha macromeris +, Coryphantha pirtlei +, Coryphantha runyonii +  and Lepidocoryphantha macromeris +
Coryphantha macromeris +
Coryphantha +
species +