Difference between revisions of "Pellaea wrightiana"

Hooker

Sp. Fil. 2: 142. 1858.

Illustrated
Synonyms: Pellaea ternifolia var. wrightiana (Hooker) A. F. Tryon
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
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|place=2: 142. 1858
 
|place=2: 142. 1858
 
|year=1858
 
|year=1858
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}}
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|special_status={{Treatment/ID/Special_status
 +
|code=F
 +
|label=Illustrated
 
}}
 
}}
 
|basionyms=
 
|basionyms=
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|name=Pellaea ternifolia var. wrightiana
 
|name=Pellaea ternifolia var. wrightiana
 
|authority=(Hooker) A. F. Tryon
 
|authority=(Hooker) A. F. Tryon
 +
|rank=variety
 
}}
 
}}
 
|hierarchy=Pteridaceae;Pellaea;Pellaea wrightiana
 
|hierarchy=Pteridaceae;Pellaea;Pellaea wrightiana
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem fragility or size;stem diameter"><b>Stems </b>compact, ascending, stout, 5–10 mm diam.;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale shape;scale width;center coloration;center width;margin coloration;margin width;margin architecture or shape">scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf arrangement or growth form;leaf some measurement"><b>Leaves </b>monomorphic, clustered on stem, 6–40 cm;</span> <span class="statement" id="st-d0_s3" data-properties="crozier pubescence">croziers sparsely villous.</span> <span class="statement" id="st-d0_s4" data-properties="petiole coloration;petiole reflectance;petiole shape;petiole architecture;articulation line prominence"><b>Petiole </b>dark-brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade architecture or shape;blade width"><b>Blade </b>linear-oblong, 2-pinnate proximally, 1.5–5 cm wide;</span> <span class="statement" id="st-d0_s6" data-properties="rachis coloration;rachis course;rachis architecture;rachis pubescence">rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous.</span> <span class="statement" id="st-d0_s7" data-properties="pinna orientation;rachis orientation;rachis shape;ultimate segment count"><b>Pinnae </b>perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–9 ultimate segments;</span> <span class="statement" id="st-d0_s8" data-properties="costa course;costa some measurement;costa height or length or size">costae straight, 2–20 mm, usually shorter than ultimate segments.</span> <span class="statement" id="st-d0_s9" data-properties="ultimate segment shape;ultimate segment some measurement;ultimate segment texture;ultimate segment pubescence"><b>Ultimate </b>segments narrowly oblong, 5–20 mm, leathery, glabrous;</span> <span class="statement" id="st-d0_s10" data-properties="margin orientation;segment reproduction;abaxial surface count;border coloration;border shape">margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate;</span> <span class="statement" id="st-d0_s11" data-properties="apex shape">apex mucronate.</span> <span class="statement" id="st-d0_s12" data-properties="vein prominence"><b>Veins </b>of ultimate segments obscure.</span> <span class="statement" id="st-d0_s13" data-properties="spore count;gland count or density"><b>Sporangia </b>long-stalked, containing 64 spores, intermixed with sparse farina-producing glands.</span> <span class="statement" id="st-d0_s14" data-properties="sporangium architecture;2n chromosome count">2n = 116.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. <b>Leaves</b> monomorphic, clustered on stem, 6–40 cm; croziers sparsely villous. <b>Petiole</b> dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. <b>Blade</b> linear-oblong, 2-pinnate proximally, 1.5–5 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. <b>Pinnae</b> perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–9 ultimate segments; costae straight, 2–20 mm, usually shorter than ultimate segments. <b>Ultimate</b> segments narrowly oblong, 5–20 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate; apex mucronate. <b>Veins</b> of ultimate segments obscure. <b>Sporangia</b> long-stalked, containing 64 spores, intermixed with sparse farina-producing glands. <b>2n</b> = 116.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
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|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates
 
|habitat=Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates
 
|elevation=300–2900 m
 
|elevation=300–2900 m
 
|distribution=Ariz.;Colo.;N.Mex.;N.C.;Okla.;Tex.;Utah;n Mexico.
 
|distribution=Ariz.;Colo.;N.Mex.;N.C.;Okla.;Tex.;Utah;n Mexico.
|discussion=<p>W. H. Wagner Jr. (1965) suggested that Pellaea wrightiana was a fertile allotetraploid hybrid between P. truncata (as P. longimucronata) and P. ternifolia. This hypothesis has been confirmed by isozyme analyses (M. D. Windham 1988). Pellaea wrightiana is therefore treated as a distinct species rather than a variety of P. ternifolia. This tetraploid species hybridizes with P. truncata and P. ternifolia subsp. arizonica to produce sterile triploids and tetraploids with intermediate morphology and malformed spores. Pellaea wrightiana has also hybridized with P. atropurpurea to form a rare apogamous pentaploid known only from western Oklahoma.</p>
+
|discussion=<p>W. H. Wagner Jr. (1965) suggested that <i>Pellaea wrightiana</i> was a fertile allotetraploid hybrid between <i>P. truncata</i> (as P. longimucronata) and <i>P. ternifolia</i>. This hypothesis has been confirmed by isozyme analyses (M. D. Windham 1988). <i>Pellaea wrightiana</i> is therefore treated as a distinct species rather than a variety of <i>P. ternifolia</i>. This tetraploid species hybridizes with <i>P. truncata</i> and <i>P. ternifolia </i>subsp.<i> arizonica</i> to produce sterile triploids and tetraploids with intermediate morphology and malformed spores. <i>Pellaea wrightiana</i> has also hybridized with <i>P. atropurpurea</i> to form a rare apogamous pentaploid known only from western Oklahoma.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Pellaea wrightiana
 
name=Pellaea wrightiana
|author=
 
 
|authority=Hooker
 
|authority=Hooker
 
|rank=species
 
|rank=species
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|basionyms=
 
|basionyms=
 
|family=Pteridaceae
 
|family=Pteridaceae
 +
|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates
 
|habitat=Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates
 
|elevation=300–2900 m
 
|elevation=300–2900 m
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|publication title=Sp. Fil.
 
|publication title=Sp. Fil.
 
|publication year=1858
 
|publication year=1858
|special status=
+
|special status=Illustrated
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_480.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_480.xml
 
|genus=Pellaea
 
|genus=Pellaea
 
|species=Pellaea wrightiana
 
|species=Pellaea wrightiana
|2n chromosome count=116
 
|abaxial surface count=0;1/2
 
|apex shape=mucronate
 
|articulation line prominence=prominent
 
|blade architecture or shape=2-pinnate
 
|blade shape=linear-oblong
 
|blade width=1.5cm;5cm
 
|border coloration=whitish
 
|border shape=crenulate
 
|center coloration=black
 
|center width=thick
 
|costa course=straight
 
|costa height or length or size=usually shorter
 
|costa some measurement=2mm;20mm
 
|crozier pubescence=villous
 
|gland count or density=sparse
 
|leaf architecture=monomorphic
 
|leaf arrangement or growth form=clustered
 
|leaf some measurement=6cm;40cm
 
|margin architecture or shape=erose-dentate
 
|margin coloration=brown
 
|margin orientation=recurved
 
|margin width=thin
 
|petiole architecture=grooved
 
|petiole coloration=dark-brown
 
|petiole reflectance=lustrous
 
|petiole shape=flattened
 
|pinna orientation=perpendicular
 
|rachis architecture=grooved
 
|rachis coloration=brown
 
|rachis course=straight
 
|rachis orientation=ascending
 
|rachis pubescence=glabrous
 
|rachis shape=not decurrent
 
|scale coloration=bicolored
 
|scale shape=linear-subulate
 
|scale width=0.1mm;0.3mm
 
|segment reproduction=fertile
 
|sporangium architecture=long-stalked
 
|spore count=64
 
|stem architecture or arrangement=compact
 
|stem diameter=5mm;10mm
 
|stem fragility or size=stout
 
|stem orientation=ascending
 
|ultimate segment count=3;9
 
|ultimate segment pubescence=glabrous
 
|ultimate segment shape=oblong
 
|ultimate segment some measurement=5mm;20mm
 
|ultimate segment texture=leathery
 
|vein prominence=obscure
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Pellaea]]
 
-->[[Category:Treatment]][[Category:Pellaea]]

Latest revision as of 21:23, 5 November 2020

Stems compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. Leaves monomorphic, clustered on stem, 6–40 cm; croziers sparsely villous. Petiole dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. Blade linear-oblong, 2-pinnate proximally, 1.5–5 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. Pinnae perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3–9 ultimate segments; costae straight, 2–20 mm, usually shorter than ultimate segments. Ultimate segments narrowly oblong, 5–20 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate; apex mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 64 spores, intermixed with sparse farina-producing glands. 2n = 116.


Phenology: Sporulating summer–fall.
Habitat: Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates
Elevation: 300–2900 m

Distribution

V2 480-distribution-map.gif

Ariz., Colo., N.Mex., N.C., Okla., Tex., Utah, n Mexico.

Discussion

W. H. Wagner Jr. (1965) suggested that Pellaea wrightiana was a fertile allotetraploid hybrid between P. truncata (as P. longimucronata) and P. ternifolia. This hypothesis has been confirmed by isozyme analyses (M. D. Windham 1988). Pellaea wrightiana is therefore treated as a distinct species rather than a variety of P. ternifolia. This tetraploid species hybridizes with P. truncata and P. ternifolia subsp. arizonica to produce sterile triploids and tetraploids with intermediate morphology and malformed spores. Pellaea wrightiana has also hybridized with P. atropurpurea to form a rare apogamous pentaploid known only from western Oklahoma.

Selected References

None.

Lower Taxa

None.
... more about "Pellaea wrightiana"
Michael D. Windham +
Hooker +
Ariz. +, Colo. +, N.Mex. +, N.C. +, Okla. +, Tex. +, Utah +  and n Mexico. +
300–2900 m +
Cliffs and rocky slopes, on a variety of acidic to mildly basic substrates +
Sporulating summer–fall. +
Illustrated +
Pellaea ternifolia var. wrightiana +
Pellaea wrightiana +
species +