Difference between revisions of "Woodsia plummerae"

Lemmon

Bot. Gaz. 7: 6. 1882.

Common names: Plummer's cliff fern
Synonyms: Woodsia obtusa var. glandulosa D. C. Eaton & M. FaxonWoodsia obtusa var. plummerae (Lemmon) MaxonWoodsia pusilla var. glandulosa (D. C. Eaton & M. Faxon) T. M. C. Taylor
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;stem orientation;petiole base count;petiole base duration"><b>Stems </b>compact, erect to ascending, with a few persistent petiole bases of unequal lengths;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale shape;central stripe coloration;margin coloration">scales often uniformly brown but at least some bicolored with dark central stripe and pale-brown margins, narrowly lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>5–25 × 1.5–6 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole coloration;petiole architecture;petiole fragility or texture"><b>Petiole </b>reddish-brown to dark purple when mature, not articulate above base, somewhat pliable and resistant to shattering.</span> <span class="statement" id="st-d0_s4" data-properties=""><b>Blade </b>lanceolate to ovate, usually 2-pinnate proximally, densely glandular, often somewhat viscid;</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade architecture or shape;blade architecture or function or pubescence;blade coating;stalk width;tip architecture">most glandular-hairs with thick stalks and distinctly bulbous tips;</span> <span class="statement" id="st-d0_s6" data-properties="glandular-hair count;scale count;scale size or width">rachis with abundant glandular-hairs and a few narrow scales.</span> <span class="statement" id="st-d0_s7" data-properties="pinna shape;pinna shape;pinna shape;pinna length or size;pinna shape;pinna shape;apex shape;apex shape"><b>Pinnae </b>ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate;</span> <span class="statement" id="st-d0_s8" data-properties="pair count">largest pinnae with 5–11 pairs of pinnules;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial and adaxial hair architecture or function or pubescence;abaxial and adaxial hair count;abaxial and adaxial hair architecture;abaxial and adaxial scale architecture or function or pubescence;abaxial and adaxial scale count;abaxial and adaxial scale architecture">abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales.</span> <span class="statement" id="st-d0_s10" data-properties="pinnule architecture or shape;pinnule shape"><b>Pinnules </b>dentate, often shallowly lobed;</span> <span class="statement" id="st-d0_s11" data-properties="margin reflectance;margin width;margin architecture or function or pubescence;cilium count;projection count;projection architecture;projection coloration or reflectance">margins nonlustrous, thin, densely glandular, lacking cilia but with occasional 1–2-celled translucent projections.</span> <span class="statement" id="st-d0_s12" data-properties="vein tip size;vein tip prominence"><b>Vein </b>tips slightly (if at all) enlarged, barely visible adaxially.</span> <span class="statement" id="st-d0_s13" data-properties="segment width"><b>Indusia </b>of relatively broad segments;</span> <span class="statement" id="st-d0_s14" data-properties="segment length;segment shape;segment architecture or arrangement;cell architecture or shape;sporangium life cycle">segments multiseriate for most of length, often divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia.</span> <span class="statement" id="st-d0_s15" data-properties="spore some measurement;2n chromosome count"><b>Spores </b>averaging 44–50 µm. <b>2n</b> = 152.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 5–25 × 1.5–6 cm. <b>Petiole</b> reddish brown to dark purple when mature, not articulate above base, somewhat pliable and resistant to shattering. <b>Blade</b> lanceolate to ovate, usually 2-pinnate proximally, densely glandular, often somewhat viscid; most glandular hairs with thick stalks and distinctly bulbous tips; rachis with abundant glandular hairs and a few narrow scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 5–11 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins nonlustrous, thin, densely glandular, lacking cilia but with occasional 1–2-celled translucent projections. <b>Vein</b> tips slightly (if at all) enlarged, barely visible adaxially. <b>Indusia</b> of relatively broad segments; segments multiseriate for most of length, often divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. <b>Spores</b> averaging 44–50 µm. <b>2n</b> = 152.</span><!--
  
 
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|phenology=Sporulating late spring–fall.
 
|habitat=Cliffs and rocky slopes, usually on granite or volcanic substrates
 
|habitat=Cliffs and rocky slopes, usually on granite or volcanic substrates
 
|elevation=700–3100 m
 
|elevation=700–3100 m
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|basionyms=
 
|basionyms=
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating late spring–fall.
 
|habitat=Cliffs and rocky slopes, usually on granite or volcanic substrates
 
|habitat=Cliffs and rocky slopes, usually on granite or volcanic substrates
 
|elevation=700–3100 m
 
|elevation=700–3100 m
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|publication year=1882
 
|publication year=1882
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_441.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_441.xml
 
|genus=Woodsia
 
|genus=Woodsia
 
|species=Woodsia plummerae
 
|species=Woodsia plummerae
|2n chromosome count=152
 
|abaxial and adaxial hair architecture=nonglandular
 
|abaxial and adaxial hair architecture or function or pubescence=glandular
 
|abaxial and adaxial hair count=lacking
 
|abaxial and adaxial scale architecture=nonglandular
 
|abaxial and adaxial scale architecture or function or pubescence=glandular
 
|abaxial and adaxial scale count=lacking
 
|apex shape=acute;rounded
 
|blade architecture or function or pubescence=glandular
 
|blade architecture or shape=2-pinnate
 
|blade coating=viscid
 
|blade shape=lanceolate;ovate
 
|cell architecture or shape=isodiametric
 
|central stripe coloration=dark
 
|cilium count=lacking
 
|glandular-hair count=abundant
 
|leaf length=5cm;25cm
 
|leaf width=1.5cm;6cm
 
|margin architecture or function or pubescence=glandular
 
|margin coloration=pale-brown
 
|margin reflectance=nonlustrous
 
|margin width=thin
 
|pair count=5;11
 
|petiole architecture=not articulate
 
|petiole base count=few
 
|petiole base duration=persistent
 
|petiole coloration=reddish-brown;dark purple
 
|petiole fragility or texture=pliable
 
|pinna length or size=longer than wide
 
|pinna shape=attenuate;tapered;ovate-deltate;elliptic
 
|pinnule architecture or shape=dentate
 
|pinnule shape=lobed
 
|projection architecture=1-2-celled
 
|projection coloration or reflectance=translucent
 
|projection count=occasional
 
|scale coloration=bicolored;brown
 
|scale count=few
 
|scale shape=lanceolate
 
|scale size or width=narrow
 
|segment architecture or arrangement=uniseriate
 
|segment length=multiseriate
 
|segment shape=divided
 
|segment width=broad
 
|sporangium life cycle=mature
 
|spore some measurement=44um;50um
 
|stalk width=thick
 
|stem architecture or arrangement=compact
 
|stem orientation=erect;ascending
 
|tip architecture=bulbous
 
|vein tip prominence=visible
 
|vein tip size=enlarged
 
 
}}<!--
 
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-->[[Category:Treatment]][[Category:Woodsia]]
 
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Revision as of 14:20, 27 July 2019

Stems compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. Leaves 5–25 × 1.5–6 cm. Petiole reddish brown to dark purple when mature, not articulate above base, somewhat pliable and resistant to shattering. Blade lanceolate to ovate, usually 2-pinnate proximally, densely glandular, often somewhat viscid; most glandular hairs with thick stalks and distinctly bulbous tips; rachis with abundant glandular hairs and a few narrow scales. Pinnae ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 5–11 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. Pinnules dentate, often shallowly lobed; margins nonlustrous, thin, densely glandular, lacking cilia but with occasional 1–2-celled translucent projections. Vein tips slightly (if at all) enlarged, barely visible adaxially. Indusia of relatively broad segments; segments multiseriate for most of length, often divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. Spores averaging 44–50 µm. 2n = 152.


Phenology: Sporulating late spring–fall.
Habitat: Cliffs and rocky slopes, usually on granite or volcanic substrates
Elevation: 700–3100 m

Distribution

V2 441-distribution-map.gif

Ariz., Calif., Colo., N.Mex., Okla., Tex., n Mexico.

Discussion

The origin and phylogenetic affinities of the tetraploid Woodsia plummerae have not been established with certainty. The hypothesis that it arose as a hybrid between the Mexican species W. mollis (Kaulfuss) J. Smith and W. mexicana Fée (D. F. M. Brown 1964; D. B. Lellinger 1985) seems untenable in light of recent chromosome studies indicating that the latter species is also tetraploid (M. D. Windham 1993). On the basis of sporophyte morphology and spore ornamentation, W. plummerae appears most closely related to the W. mexicana complex and W. oregana. In fact, W. oregana can be difficult to separate from W. plummerae in western New Mexico and northern Arizona. Intermediate plants occurring in this region may represent stable allotetraploids resulting from hybridization between the diploid progenitors of W. plummerae and W. oregana subsp. cathcartiana. Considering the available evidence, populations of W. plummerae in the United States probably originated through autopolyploidy from a recently discovered, but as yet unnamed, Mexican diploid of similar morphology. Woodsia plummerae occasionally hybridizes with W. phillipsii to produce sterile, morphologically intermediate triploids.

Selected References

None.

Lower Taxa

None.
... more about "Woodsia plummerae"
Michael D. Windham +
Lemmon +
Plummer's cliff fern +
Ariz. +, Calif. +, Colo. +, N.Mex. +, Okla. +, Tex. +  and n Mexico. +
700–3100 m +
Cliffs and rocky slopes, usually on granite or volcanic substrates +
Sporulating late spring–fall. +
Woodsia obtusa var. glandulosa +, Woodsia obtusa var. plummerae +  and Woodsia pusilla var. glandulosa +
Woodsia plummerae +
species +