Gemmabryum caespiticium

(Hedwig) J. R. Spence

Phytologia 91: 497. 2009.

Basionym: Bryum caespiticium Hedwig Sp. Musc. Frond., 180. 1801
Synonyms: Ptychostomum imbricatulum (Müller Hal.) D. T. Holyoak & N. Pedersen
Treatment appears in FNA Volume 28. Treatment on page 131. Mentioned on page 132.

Plants large. Stems 0.1–2(–3) cm. Leaves weakly to moderately concave, 0.5–2(–3) mm; margins plane to strongly revolute, limbidium distinct to weak, of 1 or 2 rows of elongate incrassate cells; apex occasionally hyaline with age; costa short- to long-excurrent, awn brown, yellow-brown, or sometimes hyaline, smooth or rarely somewhat denticulate, 1/4–3/4 leaf length; proximal laminal cells abruptly quadrate to rectangular, 1–2:1 away from costa, 2–4:1 along costa; medial and distal cells long-hexagonal, 12–18(–20) µm wide, (3–)4–6:1. Specialized asexual reproduction rare, by deciduous brood branchlets in distal leaf axils, and rhizoidal tubers, red, spheric, 100–200 µm, cells smooth. Capsule with endostome segments pale tan, brown, red-brown, or rarely pale yellow. Spores 8–14(–16) µm.

Phenology: Capsules mature Apr–Aug (spring–summer).
Habitat: Disturbed soil, earth banks, rotten wood, rock, disturbed habitats
Elevation: low to high elevations (0-4000 m)


V28 205-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., P.E.I., Que., Sask., Yukon, Alaska, Ariz., Ark., Calif., Colo., Conn., Del., D.C., Idaho, Ill., Ind., Iowa, Kans., Ky., Md., Mass., Mich., Minn., Mo., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.Dak., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wis., Wyo., Mexico (Baja California, Hidalgo), s South America, Europe, Asia, Africa, Atlantic Islands, Pacific Islands, Australia.


The combination of small caespitose plants with ovate-lanceolate concave leaves, strong awns, and inflated subalar cells on fertile stems is diagnostic of Gemmabryum caespiticium. The last character is also a diagnostic character of sect. Cladodium of Ptychostomum and known from a few other species in other genera (for example, Bryum dyffryense Holyoak).

Selected References


Lower Taxa

... more about "Gemmabryum caespiticium"
John R. Spence +
(Hedwig) J. R. Spence +
Bryum caespiticium +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +, Wyo. +, Mexico (Baja California +, Hidalgo) +, s South America +, Europe +, Asia +, Africa +, Atlantic Islands +, Pacific Islands +  and Australia. +
low to high elevations (0-4000 m) +
Disturbed soil, earth banks, rotten wood, rock, disturbed habitats +
Capsules mature Apr–Aug (spring–summer). +
Ptychostomum imbricatulum +
Gemmabryum caespiticium +
Gemmabryum sect. Caespitibryum +
species +