Pentagramma triangularis subsp. semipallida

(J. T. Howell) Yatskievych

Amer. Fern J. 80: 16. 1990.

Basionym: Pityrogramma triangularis var. semipallida J. T. Howell Leafl. W. Bot. 9: 223. 1962
Treatment appears in FNA Volume 2.

Petiole glabrous, not viscid-glandular. Blade thin and herbaceous, sometimes thick (but not leathery), not viscid-glandular, abaxially densely white-farinose, adaxially glabrous. Distal pinnae mostly regularly lobed. Proximal basiscopic lobes of basal pinnae pinnatifid, often deeply so.

Habitat: Chaparral, pine and oak woodlands
Elevation: 100–900 m


Pentagramma triangularis subsp. semipallida, as here treated, remains heterogeneous. Diploid (based on spore size) populations occur in the foothills of the Sierra Nevada in Butte County, California. Tetraploids (based on spore size) of similar morphology are apparently restricted to Santa Barbara County, California, including the adjacent Channel Islands. The relationship between these two variants has not been studied in detail.

Selected References


Lower Taxa

George Yatskievych +  and Michael D. Windham +
(J. T. Howell) Yatskievych +
Pityrogramma triangularis var. semipallida +
100–900 m +
Chaparral, pine and oak woodlands +
Amer. Fern J. +
Pityrogramma triangularis +
Pentagramma triangularis subsp. semipallida +
Pentagramma triangularis +
subspecies +