Timmiella anomala

(Bruch & Schimper) Limpricht

Laubm. Deutschl. 1: 592. 1888,.

Basionym: Barbula anomala Bruch & Schimper Bryol. Europ. 2: 107, plate 169. 1842
Synonyms: Timmiella subanomala (Bescherelle) Brotherus Trichostomum subanomala (Bescherelle) Brotherus
Treatment appears in FNA Volume 27. Treatment on page 482.

Stems commonly branched. Leaf base narrower to little wider than the limb. Sexual condition dioicous, autoicous or synoicous, occasionally polyoicous. Seta usually 1.5–2.5 cm. Capsule with theca reaching 3–4 mm, operculum commonly 1–1.5 mm, peristome distinctly twisted.


Phenology: Sporophytes mature summer–fall (occasionally winter).
Habitat: Soil, humus, limestone crevices, vertical rock face
Elevation: moderate elevations (700-1500 m)

Distribution

V27 697-distribution-map.gif

Ariz., Calif., N.Mex., Tex., Wash., Mexico, Central America, Europe, Asia, n Africa.

Discussion

Timmiella anomala commonly grows in dusty, friable soil that makes dissections difficult. H. A. Crum and L. E. Anderson (1980–1983, vol. 4) suggested that this species may also be dioicous; K. Saito (1975) and Li X. J. et al. (2001) asserted that it is dioicous in Asia (Japan and China, respectively). A specimen from California, Showers 3326, MO, has stems with synoicous perichaetia and also apparently entirely perigoniate stems, possibly due to rhizautoicy. According to B. Allen (2002), Timmiella anomala in Central America has peristome teeth erect to very weakly twisted though these are described as at least once twisted in publications on other areas of the world. Material from the Yukon is sterile and only tentatively identifiable as this species.

Selected References

None.

Lower Taxa

None.
... more about "Timmiella anomala"
Richard H. Zander +
(Bruch & Schimper) Limpricht +
Barbula anomala +
Ariz. +, Calif. +, N.Mex. +, Tex. +, Wash. +, Mexico +, Central America +, Europe +, Asia +  and n Africa. +
moderate elevations (700-1500 m) +
Soil, humus, limestone crevices, vertical rock face +
Sporophytes mature summer–fall (occasionally winter). +
Laubm. Deutschl. +
Timmiella subanomala +  and Trichostomum subanomala +
Timmiella anomala +
Timmiella +
species +