Tortula protobryoides

R. H. Zander

Bull. Buffalo Soc. Nat. Sci. 32: 226. 1993,.

Basionym: Phascum bryoides Dickson Fasc. Pl. Crypt. Brit. 4: 3, plate 10, fig. 3. 1801,
Synonyms: Pottia bryoides (Dickson) Mitten Protobryum bryoides (Dickson) J. Guerra & M. J. Cano
Treatment appears in FNA Volume 27. Treatment on page 592. Mentioned on page 587, 589, 629.

Leaves ovate to oblong-lanceolate, apex broadly acute to rounded, mucronate to short-awned, margins revolute in mid 2/3–3/4 of leaf, weakly bordered from apex to near base with (2–)4 rows of weakly thicker-walled and less-papillose or smooth cells; costa excurrent, lacking an adaxial pad of cells though convex adaxially, distally narrow, 3–4 cells across adaxial surface; distal laminal cells subquadrate to hexagonal, occasionally somewhat elongated longitudinally, width 13–18 µm wide, 1(–2):1, papillose with 4(–6) simple papillae per lumen. Sexual condition autoicous. Sporophytes exerted. Seta length 0.2–0.4 cm. Capsule stegocarpic, not systylius, broadly cylindric, erect and nearly straight, urn ca. 1.4–1.5 mm; peristome remaining adherent to the inside of the operculum, teeth rudimentary to linear, to 300 µm, of several filaments variously anastomosing and weakly twisted, basal membrane length to 50 µm; operculum ca. 0.7–0.8 mm, not falling. Spores ca. 20–25 µm, spheric, densely low-papillose.

Phenology: Capsules mature winter–spring.
Habitat: Soil
Elevation: low elevations


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B.C., Sask., Ariz., Calif., Colo., Minn., Mexico (Baja California), Europe, sw Asia.


Selected References


Lower Taxa

... more about "Tortula protobryoides"
Richard H. Zander +  and Patricia M. Eckel +
R. H. Zander +
Phascum bryoides +
B.C. +, Sask. +, Ariz. +, Calif. +, Colo. +, Minn. +, Mexico (Baja California) +, Europe +  and sw Asia. +
low elevations +
Capsules mature winter–spring. +
Bull. Buffalo Soc. Nat. Sci. +
Pottia bryoides +  and Protobryum bryoides +
Tortula protobryoides +
species +